← Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, First Part: Porulosa (Spumellaria and Acantharia) / Report on the Scientific Results of the Voyage of H.M.S. Challen

Subfamily 1. Cenolarcida, Haeckel

Definition.—Larcarida with a simple, lentelliptical latticed shell (cortical shell without a medullary shell).

Genus 266. Cenolarcus, n. gen.

Definition.—Larcarida with a simple, lentelliptical latticed shell, without a medullary shell, without radial spines.

The genus Cenolarcus begins the group of Larcoidea as the most simple form of this suborder. It corresponds to Cenosphæra among the Sphæroidea, to Cenodiscus among the Discoidea, to Cenellipsis among the Prunoidea. The simple latticed shell is distinguished from that of the three other genera by its typical lentelliptical form, a triaxial ellipsoid with three dimensive axes of unequal length. Probably Cenolarcus is the original ancestral form of the Larcoidea, derived from Actilarcus (the lentelliptical Actissa) by the formation of a simple fenestrated shell around the lentelliptical central capsule. But possibly also some species of Cenolarcus may be derived from Coccolarcus or Larnacilla by reduction and loss of the medullary shell (compare Cenolarcus triaxonius, n. sp.).

1. Cenolarcus primordialis, n. sp. (Pl. , figs. 7, 7a, 7b).

Network of the shell regular, with circular, hexagonally framed pores, twice as broad as the elevated bars; about ten pores on the half meridian, eight on the half equator. Surface a little rough. Proportion of the three dimensive axes = 2 : 2.5 : 3.

Dimensions.—Principal axis (or length) 0.12, transverse axis (or breadth) 0.1, sagittal axis (or thickness) 0.08; pores 0.01, bars 0.005.

Habitat.—Pacific, central area, Station 271, depth 2425 fathoms.

2. Cenolarcus dimensivus, n. sp.

Network of the shell regular, with circular pores (without hexagonal frames), three times as broad as the thin bars; about twelve pores on the half meridian, nine on the half equator. Surface thorny. Proportion of the three dimensive axes = 3 : 4 : 5.

Dimensions.—Principal axis 0.15, transverse axis 0.12, sagittal axis 0.09; pores 0.012, bars 0.004.

Habitat.—North Pacific, Station 244, surface.

3. Cenolarcus triaxonius, n. sp.

Network of the shell regular, with circular, hexagonally framed pores, four times as broad as the thin bars; about twelve pores on the half meridian, nine on the equator. Surface smooth. From the inner surface of the shell arise six very thin radial beams, opposite in pairs in the three dimensive axes; all six beams end freely in a little knob, at an equal distance from the centre; therefore this remarkable species seems to have lost a medullary shell (descending from Coccolarcus or Larnacilla?). Proportion of the three dimensive axes = 2 : 2.5 : 3.

Dimensions.—Principal axis 0.13, transverse axis 0.11, sagittal axis 0.09; pores 0.012, bars 0.03.

Habitat.—Tropical Atlantic, Station 338, depth 1990 fathoms.

4. Cenolarcus lentellipticus, n. sp.

Network of the shell regular, with circular pores of the same breadth as the thick bars; about eighteen pores on the half meridian, fourteen on the half equator. Surface smooth. Proportion of the three dimensive axes = 2 : 3 : 4.

Dimensions.—Principal axis 0.16, transverse axis 0.12, sagittal axis 0.08; pores and bars 0.004.

Habitat.—Western Tropical Pacific, Station 224, surface.

5. Cenolarcus minimus, n. sp.

Network of the shell subregular, with very small circular pores of the same breadth as the bars; only four pores on the half meridian, three on the half equator. Surface smooth. Proportion of the three dimensive axes = 3 : 4 : 5.

Dimensions.—Principal axis 0.05, transverse 0.04, sagittal axis 0.03; pores and bars 0.006.

Habitat.—Pacific, central area, Station 266, depth 2750 fathoms.

Genus 267. Larcarium, n. gen.

Definition.—Larcarida with a simple, lentelliptical latticed shell, without a medullary shell; surface covered with radial spines.

The genus Larcarium differs from Cenolarcus only in the possession of radial spines on the surface of the simple fenestrated lentelliptical shell. These spines are commonly disposed symmetrically, opposite in pairs, either on the poles of the three dimensive axes, or on the poles of certain diagonal axes. Larcarium differs from the similar genera Larcidium, Larnacidium, and Larnacantha by the absence of any medullary shell.

1. Larcarium amphistylum, n. sp.

Shell thorny, with two large conical spines, opposite on both poles of the principal axis, somewhat longer than it. Network of the shell regular, with circular, hexagonally framed pores, twice as broad as the bars; about twelve pores on the half meridian, ten on the half equator. Proportion of the three dimensive axes = 3 : 4 : 5.

Dimensions.—Principal axis (or length) 0.15, transverse axis (or breadth) 0.12, sagittal axis (or thickness) 0.09; pores 0.01, bars 0.005.

Habitat.—Pacific, central area, Station 274, depth 2750 fathoms.

2. Larcarium staurostylum, n. sp.

Shell smooth, with four short three-sided pyramidal spines of equal length, opposite in pairs on the poles of the principal and lateral axes. Network of the shell regular, with circular pores of the same breadth as the bars; about eight pores on the half meridian, six on the half equator. Proportion of the three dimensive axes = 2 : 2.5 : 3.

Dimensions.—Length of the shell 0.12, breadth 0.1, thickness 0.08; pores and bars 0.006.

Habitat.—South Pacific, Station 300, depth 1375 fathoms.

3. Larcarium hexastylum, n. sp.

Shell smooth, with six short conical spines of equal length (= the shortest axis of the shell), opposite in pairs on the poles of the three dimensive axes. Network of the shell subregular, with circular pores three times as broad as the bars; about eleven pores on the half meridian, nine on the half equator. Proportion of the three dimensive axes = 3 : 4 : 5.

Dimensions.—Length of the shell 0.1, breadth 0.08, thickness 0.06, pores 0.006; bars 0.002.

Habitat.—North Pacific, Station 244, surface.

4. Larcarium axostylum, n. sp.

Shell thorny, with six strong conical radial spines, opposite in pairs on the poles of the three dimensive axes. All three pairs are of different sizes, the length of each spine corresponding nearly to the size of the shell-axis, of which it is the prolongation. Network of the shell subregular, with circular pores twice as broad as the bars; ten on the half meridian, eight on the half equator. Proportion of the three dimensive axes = 2 : 3 : 4.

Dimensions.—Length of the shell 0.12, breadth 0.09, thickness 0.06; pores 0.01, bars 0.005.

Habitat.—Western Tropical Pacific, Station 225, depth 4475 fathoms.

5. Larcarium octostylum, n. sp.

Shell thorny, with eight thin cylindrical radial spines, opposite in pairs in two crossed diagonal planes. Network of the shell irregular, with roundish pores, twice to four times as broad as the bars; nine to eleven on the half meridian, five to seven on the half equator. Proportion of the three dimensive axes = 3 : 4 : 5.

Dimensions.—Length of the shell 0.11, breadth 0.09, thickness 0.07; pores 0.006 to 0.012, bars 0.003.

Habitat.—Pacific, central area, Station 267, surface.

6. Larcarium polystylum, n. sp.

Shell thorny, with numerous (twenty to thirty) stronger conical radial spines, about as long as the shortest axis of the shell. Network of the shell irregular, with roundish pores, twice to four times as broad as the bars; seven to eight on the half meridian, five to six on the half equator. Proportion of the three dimensive axes = 1 : 2 : 3.

Dimensions.—Length of the shell 0.09, breadth 0.06, thickness 0.03; pores 0.006 to 0.012, bars 0.003.

Habitat.—South Atlantic, Station 323, surface.

7. Larcarium chætostylum, n. sp.

Shell bristly, with very numerous (sixty to eighty or more) very thin, bristle-like, radial spines, somewhat longer than the longest axis of the shell. Network irregular, with very small roundish pores, about the same size as the bars; sixteen to eighteen on the half meridian, thirteen to fifteen on the half equator. Proportion of the three dimensive axes = 1 : 1.5 : 2.

Dimensions.—Length of the shell (without spines) 0.13, breadth 0.1, thickness 0.07; pores and bars 0.004 to 0.006.

Habitat.—Pacific, central area, Station 273, surface.