Division II. Dicotyledons (part 2)

Alnus inhabits swamps, river bottom-lands, and high mountains, and is widely and generally distributed through the northern hemisphere, often forming the most conspicuous feature of vegetation on mountain slopes, ranging at high altitudes southward in the New World through Central America to Colombia, Peru, and Bolivia, and to upper Assam and Japan in the Old World. Of the eighteen or twenty species now recognized nine are North American; of these, six attain the size and habit of trees. Of the exotic species, Alnus vulgaris Hill., a common European, north African, and Asiatic timber-tree, was introduced many years ago into the northeastern states, where it has become locally naturalized. The wood of Alnus is very durable in water, and the astringent bark and strobiles are used in tanning leather and in medicine.

Alnus is the classical name of the Alder.

Conspectus of the North American Arborescent Species.

Flowers opening in spring with or after the leaves; stamens 4; pistillate aments inclosed during the winter; wing of the nut broad; leaves ovate, sinuately lobed, lustrous on the lower surface.

1. A. sinuata (B, F, G).

Flowers opening in winter or early spring before the unfolding of the leaves; pistillate aments usually naked during the winter.

Wing of the nut broad; leaves ovate or elliptic, rusty-pubescent on the lower surface; pistillate aments often inclosed during the winter; stamens 4.

2. A. rubra (B, G).

Wing of the nut reduced to a narrow border.

Stamens 4; leaves oblong-ovate, glabrous or puberulous on the lower surface.

3. A. tenuifolia (B, F, G).

Stamens usually 2, or 3.

Leaves ovate or oval.

4. A. rhombifolia (B, F, G).

Leaves oblong-lanceolate, acute.

5. A. oblongifolia (H).

Flowers opening in autumn from aments of the year; stamens 4; wing of the nut reduced to a narrow border; leaves oblong-ovate or obovate, dark green and lustrous above, pale yellow-green below.

6. A. maritima (A).

1. Alnus sinuata Rydb. Alder.

Alnus sitchensis Sarg.

Leaves ovate, acute, full and rounded and often unsymmetrical and somewhat oblique or abruptly narrowed and cuneate at base, divided into numerous short acute lateral lobes, sharply and doubly serrate with straight glandular teeth, glandular-viscid as they unfold, at maturity membranaceous, yellow-green on the upper surface, pale and very lustrous on the lower surface, glabrous, or villose along the under side of the stout midrib with short brown hairs also forming tufts in the axils of the numerous slender primary veins, 3′—6′ long, 1½′—4′ wide; petioles stout, grooved, abruptly enlarged at the base, ½′—¾′ in length; stipules oblong to spatulate, rounded and apiculate at apex, puberulous, about ¼′ long. Flowers: staminate aments sessile, in pairs in the axils of the upper leaves sometimes reduced to small bracts, and single in the axil of the leaf next below, during the winter about ½′ long and ⅛′ thick, with dark red-brown shining puberulous apiculate scales, becoming when the flowers open from spring to midsummer 4′ or 5′ long, with a puberulous light red rachis and ovate acute apiculate 3-flowered scales; calyx-lobes rounded, shorter than the 4 stamens; pistillate aments in elongated panicles, inclosed during winter in buds formed the previous summer in the axils of the leaves of short lateral branchlets, long-pedunculate, ⅓′ long, ⅙′ thick. Fruit: strobiles on slender peduncles in elongated sometimes leafy panicles 4′—6′ in length, oblong, ½′—⅝′ long, about ⅓′ thick, their truncate scales thickened at the apex; nut oval, about as wide as its wings.

A tree, sometimes 40° high, with a trunk 7′—8′ in diameter, short small nearly horizontal branches forming a narrow crown, and slender slightly zigzag branchlets puberulous and very glandular when they first appear, bright orange-brown and lustrous and marked by numerous large pale lenticels during their first season, much roughened during their second year by the elevated crowded leaf-scars, becoming light gray. Winter-buds acuminate, dark purple, covered especially toward the apex with close fine pubescence, about ½′ long. Bark thin bluish gray, with bright red inner bark; often a shrub only a few feet tall spreading into broad thickets.

Distribution. Northwest coast from the borders of the Arctic Circle to the high mountains of northern California; common in the valley of the Yukon and eastward through British Columbia to Alberta, and through Washington and Oregon to the western slopes of the Rocky Mountains in Montana; at the north with dwarf Willows, forming great thickets; in southeastern Alaska often a tall tree on rich moist bottom-lands near the mouths of mountain streams, and at the upper limits of tree growth a low shrub; very abundant in the valley of the Yukon on the wet banks of streams and often arborescent in habit; in British Columbia and the United States generally smaller and a shrub, growing usually only at altitudes of more than 3000° above the sea, and often forming thickets on the banks of streams and lakes.

2. Alnus rubra Bong. Alder.

Alnus oregona Nutt.

Leaves ovate to elliptic, acute, abruptly or gradually narrowed and cuneate at base, crenately lobed, dentate with minute gland-tipped teeth, and slightly revolute on the margins, covered when they unfold with pale tomentum, at maturity thick dark green and glabrous or pilose with scattered white hairs above, clothed below with short rusty pubescence, 3′—5′ long, 1¾′—3′ wide, or on vigorous branchlets sometimes 8′—10′ long, with a broad midrib and primary veins green on the upper side and orange-colored on the lower, the primary veins running obliquely to the points of the lobes and connected by conspicuous slightly reticulate cross veinlets; petioles orange-colored, nearly terete, slightly grooved, ¼′—¾′ in length; stipules ovate, acute, pale green flushed with red, tomentose, ⅛′—¼′ long. Flowers: staminate aments in red-stemmed clusters, during the winter 1¼′ long, ⅛′ thick, with dark red-brown lustrous closely appressed scales, becoming 4′—6′ long and ¼′ thick, with ovate acute orange-colored glabrous scales; calyx yellow, with ovate rounded lobes rather shorter than the 4 stamens; pistillate aments in short racemes usually inclosed during the winter in buds formed during the early summer and opening in the early spring, ⅓′—½′ long, about /16′ thick, with dark red acute scales; styles bright red. Fruit: strobiles raised on stout orange-colored peduncles sometimes ½′ in length, ovoid or oblong, ½′—1′ long, ⅓′—½′ thick, with truncate scales much thickened toward the apex; nut orbicular to obovoid, surrounded by a membranaceous wing.

A tree, usually 40°—50°, occasionally 90° high, with a trunk sometimes 3° in diameter, slender somewhat pendulous branches forming a narrow pyramidal head, and slender branchlets marked by minute scattered pale lenticels, light green and coated at first with hoary tomentum sometimes persistent until their second year, becoming during the first winter bright red and lustrous and ultimately ashy gray. Winter-buds about ⅓′ long, dark red, covered with pale scurfy pubescence. Bark rarely more than ¾′ thick, close, roughened by minute wart-like excrescences, pale gray or nearly white, with a thin outer layer, and bright red-brown inner bark. Wood light, soft, brittle, not strong, close-grained, light brown tinged with red, with thick nearly white sapwood; in Washington and Oregon largely used in the manufacture of furniture and for smoking salmon; by the Indians of Alaska the trunks are hollowed into canoes.

Distribution. Shores of Yakutat Bay, southeastern Alaska, southward near the coast to the cañons of the Santa Inez Mountains, Santa Barbara County, California; common along the banks of streams, and of its largest size near the shores of Puget Sound; in California most abundant in Mendocino, Humboldt and Marin Counties, forming groves on bottom-lands near the coast; often ranging inland for 20 or 30 miles, and occasionally ascending to altitudes of 2000° above the sea.

3. Alnus tenuifolia Nutt. Alder.

Leaves ovate-oblong, acute or acuminate, broad and rounded or cordate or occasionally abruptly narrowed and cuneate at base, usually acutely laciniately lobed and doubly serrate, when they unfold light green often tinged with red, pilose on the upper surface and coated on the lower with pale tomentum, at maturity thin and firm, dark green and glabrous above, pale yellow-green and glabrous or puberulous below, 2′—4′ long, 1½′—2½′ wide, with a stout orange-colored midrib impressed on the upper side, and slender primary veins running to the points of the lobes; petioles stout, slightly grooved, orange-colored, ½′—1′ in length; stipules ovate, acute, thin, and scarious, ½′ long, about ⅛′ wide, covered with pale pubescence. Flowers: staminate aments 3 or 4 in number in slender-stemmed racemes, nearly sessile or raised on stout peduncles often ½′ long, during the winter light purple, ¾′—1′ long and ¼′ thick, becoming 1½′—2′ in length; calyx-lobes rounded, shorter than the 4 stamens; pistillate aments naked during the winter, dark red-brown, nearly ¼′ long, with acute apiculate loosely imbricated scales, only slightly enlarged when the flowers open. Fruit: strobiles obovoid-oblong, ⅓′—½′ long, their scales much thickened, truncate and 3-lobed at apex; nut nearly circular to slightly obovoid, surrounded by a thin membranaceous border.

A tree, occasionally 30° tall, with a trunk 6′—8′ in diameter, small spreading slightly pendulous branches forming a narrow round-topped head, and slender branchlets marked at first by a few large orange-colored lenticels and coated with fine pale or rusty caducous pubescence, becoming light brown or ashy gray more or less deeply flushed with red in their first winter and ultimately paler; more often shrubby, with several spreading stems, and at the north and at high altitudes frequently only 4°—5° tall. Winter-buds ¼′—⅓′ long, bright red, and puberulous. Bark rarely more than ¼′ thick, bright red-brown and broken on the surface into small closely appressed scales.

Distribution. Banks of streams and mountain cañons from Francis Lake in latitude 61° north to the valley of the lower Fraser River, British Columbia, eastward along the Saskatchewan to Prince Albert, and southward through the Rocky Mountains to northern New Mexico; on the Sierra Nevada of southern California, and in Lower California; the common Alder of mountain streams in the northern interior region of the continent; very abundant on the eastern slopes of the Cascade Mountains, and on the southern California Sierras; forming great thickets at 6000°—7000° above the sea along the headwaters of the rivers of southern California flowing to the Pacific Ocean; the common Alder of eastern Washington and Oregon, and of Idaho and Montana; very abundant and of its largest size in Colorado and northern New Mexico.

4. Alnus rhombifolia Nutt. White Alder. Alder.

Leaves ovate or oval or sometimes nearly orbicular, rounded or acute at apex, especially on vigorous shoots, gradually or abruptly narrowed and cuneate at base, finely or sometimes coarsely and occasionally doubly serrate, slightly thickened and reflexed on the somewhat undulate margins, when they unfold pale green and covered with deciduous matted white hairs, at maturity dark green and lustrous on the upper surface, frequently marked, especially on the midrib, with minute glandular dots, light yellow-green and slightly puberulous below, 2′—3′ long, 1½′—2′ wide, with a stout yellow midrib and primary veins; petioles slender, yellow, hairy, flattened and grooved on the upper side, ½′—¾′ long; stipules ovate, acute, scarious, puberulous, about ¼′ in length. Flowers: staminate aments in slender-stemmed pubescent clusters, usually short-stalked, during the summer dark olive-brown and lustrous, ¾′—1′ long and about /16′ thick, beginning to lengthen late in the autumn before the leaves fall, fully grown and 4′—6′ long and ¼′ thick in January, with dark orange-brown scales, and deciduous in February before the appearance of the new leaves; calyx yellow, 4-lobed, rather shorter than the 2 or occasionally 3 or rarely single stamen; pistillate aments in short pubescent racemes emerging from the bud in December, their scales broadly ovate and rounded. Fruit: strobiles oblong, ⅓′—½′ long, with thin scales slightly thickened and lobed at apex, fully grown at midsummer, remaining closed until the trees flower the following year; nut broadly ovoid, with a thin margin.

A tree, frequently 70°—80° high, with a tall straight trunk 2°—3° in diameter, long slender branches pendulous at the ends, forming a wide round-topped open head, and slender branchlets marked by small scattered lenticels, at first light green and coated with pale caducous pubescence, soon becoming dark orange-red and glabrous, and darker during the winter and following summer. Winter-buds nearly ½′ long, very slender, dark red, and covered with pale scurfy pubescence. Bark on old trunks 1′ thick, dark brown, irregularly divided into flat often connected ridges broken into oblong plates covered with small closely appressed scales. Wood light, soft, not strong, brittle, close-grained, light brown, with thick lighter colored often nearly white sapwood.

Distribution. Banks of streams from northern Idaho to the eastern slope of the Cascade Mountains of Washington and southeastern Oregon, and southward from the valley of the Willamette River, Oregon (near Salem, Marion County, J. C. Nelson) over the coast ranges and along the western slopes of the Sierra Nevada to the mountains of southern California (San Bernardino, San Jacinto, and Cuyamaca Ranges); the common Alder of the valleys of central California, occasionally ascending on the southern Sierra Nevada to altitudes of 8000°, and the only species at low altitudes in the southern part of the state.

5. Alnus oblongifolia Torr. Alder.

Alnus acuminata Sarg., not H. B. K.

Leaves oblong-lanceolate, acute; or rarely obovate and rounded at apex, gradually narrowed and cuneate at base, sharply and usually doubly serrate, more or less thickly covered, especially early in the season, with black glands, dark yellow-green and glabrous or slightly puberulous above, pale and glabrous or puberulous below, especially along the slender yellow midrib and veins, with small tufts of rusty hairs in the axils of the primary veins, 2′—3′ long, about 1½′ wide; petioles slender, grooved, pubescent, ¾′ long; stipules ovate-lanceolate, brown and scarious, about ¼′ in length. Flowers: staminate aments in short stout-stemmed racemes, during the winter light yellow, ½′—¾′ long and about /16′ thick, becoming when the flowers open at the end of February before the appearance of the leaves 2′—2½′ in length, with ovate pointed dark orange-brown scales; calyx 4-lobed; stamens 3 or occasionally 2, with pale red anthers soon becoming light yellow; pistillate aments naked during the winter, ⅛′ to nearly ¼′ long, with light brown ovate rounded scales; stigmas bright red. Fruit: strobiles ½′—1′ long, with thin scales slightly thickened and nearly truncate at apex; nut broadly ovoid, with a narrow membranaceous border.

A tree, in the United States rarely more than 20°—30° high, with a trunk sometimes 8′ in diameter, long slender spreading branches forming an open round-topped head, and slender branchlets slightly puberulous when they first appear, light orange-red and lustrous during their first winter, and marked by small conspicuous pale lenticels, becoming in their second year dark red-brown or gray tinged with red and much roughened by the elevated leaf-scars. Winter-buds acute, red, lustrous, glabrous, ½′ long. Bark thin, smooth, light brown tinged with red.

Distribution. Banks of streams in cañons of the mountains of southern New Mexico and Arizona at altitudes of 4000°—6000° above the sea; in Oak Creek Cañon near Flagstaff, northern Arizona (tree 100° × 3°, P. Lowell); and on the mountains of northern Mexico.

6. Alnus maritima Nutt. Alder.

Leaves oblong-ovate, or obovate, acute, acuminate or rounded at apex, gradually narrowed and cuneate at base, remotely serrate with minute incurved glandular teeth, and somewhat thickened on the slightly undulate margins, when they unfold, light green tinged with red, hairy on the midrib, veins, and petioles, and coated above with pale scurfy pubescence, at maturity dark green, very lustrous, and covered below by minute pale glandular dots, 3′—4′ long, 1½′—2′ wide, with a stout yellow midrib and primary veins prominent and glandular on the upper side and slightly puberulous below; petioles stout, yellow, glandular, flattened and grooved on the upper side, ½′—¾′ in length; stipules oblong, acute, about ⅛′ long, dark reddish brown, caducous. Flowers opening in the autumn: aments appearing in July on branches of the year and fully grown in August or early in September; staminate in short scurfy-pubescent glandular-pitted racemes on slender peduncles sometimes 1′ in length from the axils of upper leaves, covered at first with ovate acute dark green very lustrous scales slightly ciliate on the margins and furnished at apex with minute red points, at maturity 1½′—2½′ long, ¼′ to nearly ½′ thick, with dark orange-brown scales raised on slender stalks, and 4 bright orange-colored stamens; pistillate usually solitary from the axils of the lower leaves on stout pubescent peduncles, bright red at apex and light green below before opening, with ovate acute scales slightly ciliate on the margins, about ⅛′ long when the styles protrude from between the scales, beginning to enlarge the following spring. Fruit attaining full size at midsummer and then raised on a stout peduncle, broadly ovoid, rounded and depressed at base, gradually narrowed to the rather obtuse apex, about ⅝′ long and ½′ broad, with thin lustrous scales slightly thickened and crenately lobed at apex, turning dark reddish brown or nearly black and opening late in the autumn and remaining on the branches until after the flowers open the following year; nut oblong-obovoid, gradually narrowed and apiculate at apex, with a thin membranaceous border.

A tree, occasionally 30° high, with a tall straight trunk 4′—5′ in diameter, small spreading branches forming a narrow round-topped head, slender slightly zigzag branchlets, light green and hairy at first, pale yellow-green, very lustrous, slightly puberulous, marked with occasional small orange-colored lenticels, and glandular with minute dark glandular dots during their first summer, becoming dull light orange or reddish brown in the winter, and ashy gray often slightly tinged with red the following season; more often shrubby, with numerous slender spreading stems 15°—20° tall. Winter-buds acute, dark red, coated with pale lustrous scurfy pubescence, about ¼′ long. Bark ⅛′ thick, smooth, light brown or brown tinged with gray. Wood light, soft, close-grained, light brown, with thick hardly distinguishable sapwood.

Distribution. Banks of streams and ponds in southern Delaware and Maryland, and in south central Oklahoma (Johnson and Bryan Counties).

Occasionally cultivated as an ornamental tree in the eastern states and hardy as far north as Massachusetts.

X. Fagaceæ.

Trees, with watery juice, slender terete branchlets marked by numerous usually pale lenticels, alternate stalked penniveined leaves, and narrow mostly deciduous stipules. Flowers monœcious, the staminate in unisexual heads or aments, composed of a 4—8-lobed calyx, and 4 or 8 stamens, with free simple filaments and introrse 2-celled anthers, the cells parallel and contiguous, opening longitudinally; the pistillate solitary or clustered, in terminal unisexual or bisexual spikes or heads, subtended by an involucre of imbricated bracts becoming woody and partly or entirely inclosing the fruit, and composed of a 4—8-lobed calyx adnate to the 3—7-celled ovary with as many styles as its cells and 1 or 2 pendulous anatropous or semianatropous ovules in each cell. Fruit a nut 1-seeded by abortion, the outer coat cartilaginous, the inner membranaceous or bony. Seed filling the cavity of the nut, without albumen; seed-coat membranaceous; cotyledons fleshy, including the minute superior radicle; hilum, basal, minute.

The six genera of this widely distributed family occur in North America with the exception of Nothofagus, separated from Fagus to receive the Beech-trees of the southern hemisphere.

Conspectus of the North American Genera.

Staminate flowers fascicled in globose-stalked heads; the pistillate in 2—4-flowered clusters.

1. Fagus.

Staminate flowers in slender aments.

Pistillate flowers in 2—5-flowered clusters below the staminate, in bisexual aments.

Nut inclosed in a prickly burr.

Leaves deciduous; ovary 6-celled; nut maturing in one season; branchlets lengthening by an upper axillary bud; bud-scales 4.

2. Castanea.

Leaves persistent; ovary 3-celled; nut maturing at the end of the second season; branchlets lengthening by a terminal bud; bud-scales numerous.

3. Castanopsis.

Nut inclosed only partly in a shallow cup covered by slender recurved scales united only at the base, free above.

4. Lithocarpus.

Pistillate flowers solitary, in few-flowered unisexual spikes; nut more or less inclosed in a cup covered by thin or thickened scales, closely appressed or often free toward its rim.

5. Quercus.

1. Fagus L. Beech.

Trees, with smooth pale bark, hard close-grained wood, and elongated acute bright chestnut-brown buds, their inner scales accrescent and marking the base of the branchlets with persistent ring-like scars. Leaves convex and plicate along the veins in the bud, thick and firm, deciduous; petioles short, nearly terete, in falling leaving small elevated semioval leaf-scars, with marginal rows of minute fibro-vascular bundle-scars; stipules linear-lanceolate, infolding the leaf in the bud. Flowers vernal after the unfolding of the leaves; staminate short-pedicellate, in globose many-flowered heads on long drooping bibracteolate stems at base of shoots of the year or from the axils of their lowest leaves, and composed of a subcampanulate 4—8-lobed calyx, the lobes imbricated in æstivation, ovate and rounded, and 8—16 stamens inserted on the base of and longer than the calyx, with slender filaments and oblong green anthers; pistillate in 2—4-flowered stalked clusters in the axils of upper leaves of the year, surrounded by numerous awl-shaped hairy bracts, the outer bright red, longer than the flowers, deciduous, the inner shorter and united below into a 4-lobed involucre becoming at maturity woody, ovoid, thick-walled, and covered by stout recurved prickles, inclosing or partly inclosing the usually 3 nuts, and ultimately separating into 4 valves; calyx urn-shaped, villose, divided into 4 or 5 linear-lanceolate acute lobes, its 3-angled tube adnate to the 3-celled ovary surmounted by 3 slender recurved pilose styles green and stigmatic toward the apex and longer than the involucre; ovules 2 in each cell. Nut ovoid, unequally 3-angled, acute or winged at the angles, concave and longitudinally ridged on the sides, chestnut-brown and lustrous, tipped with the remnants of the styles, marked at the base by a small triangular scar, with a thin shell covered on the inner surface with rufous tomentum. Seed dark chestnut-brown, suspended with the abortive ovules from the tip of the hairy dissepiment of the ovary pushed by the growth of the seed into one of the angles of the nut; cotyledons sweet, oily, plano-convex.

Fagus as here limited is confined to the northern hemisphere, with a single American species and seven Old World species; of these one is widely distributed through Europe, another is found in the Caucasus, and the others are confined to eastern temperate Asia. Of exotic species, the European Fagus sylvatica L., an important timber-tree, is frequently planted for ornament in the eastern states in several of its forms, especially those with purple leaves, and with pendulous branches. The wood of Fagus is hard and close-grained. The sweet seeds are a favorite food of swine, and yield a valuable oil.

Fagus is the classical name of the Beech-tree.

1. Fagus grandifolia Ehrh. Beech.

Fagus americana Sweet.

Leaves remote at the ends of the branches and clustered on short lateral branchlets, oblong-ovate, acuminate with a long slender point, coarsely serrate with spreading or incurved triangular teeth except at the gradually narrowed generally cuneate base, when they unfold pale green and clothed on the lower surface and margins with long pale lustrous silky hairs, at maturity dull dark bluish green above, light yellow-green, very lustrous, and glabrous or rarely pilose below (f. pubescens Fern. & Rehd.) with tufts of long pale hairs in the axils of the veins, 2½′—5′ long, 1′—3′ wide, with a slender yellow midrib covered above with short pale hairs, and slender primary veins running obliquely to the points of the teeth; turning bright clear yellow in the autumn; very rarely deeply laciniate; petioles hairy, ¼′—½′ length; stipules ovate-lanceolate on the lower leaves, strap-shaped to linear-lanceolate on the upper, brown or often red below the middle, membranaceous, lustrous, 1′—1½′ long. Flowers opening when the leaves are about one third grown; staminate in globose heads 1′ in diameter, on slender hairy peduncles about 2′ long; pistillate in usually 2-flowered clusters, on short clavate hoary peduncles ½′—¾′ long. Fruit: involucres ½′—¾′ in length often shorter than the nuts, on stout hairy club-shaped peduncles ¼′—¾′ long, fully grown at midsummer, and then puberulous, dark orange-green, and covered by long slender recurved prickles red above the middle, becoming at maturity in the autumn light brown and tomentose, with crowded much recurved pubescent prickles, persistent on the branch after opening late into the winter; nut about ¾′ long.

A tree, usually 70°—80° but exceptionally 120° high, sending up from the roots numerous small stems sometimes extending into broad thickets round the parent tree, in the forest with a long comparatively slender stem free of branches for more than half its length, and short branches forming a narrow head, in open situations short-stemmed, with a trunk often 3°—4° in diameter, and numerous limbs spreading gradually and forming a broad compact round-topped head of slender slightly drooping branches clothed with short leafy laterals, and branchlets pale green and coated with long soft caducous hairs when they first appear, olive-green or orange-colored during their first summer, and conspicuously marked by oblong bright orange lenticels, gradually growing red, bright reddish brown during their first winter, darker brown in their second season and ultimately ashy gray. Winter-buds puberulous, especially toward the apex, ¾′ to nearly 1′ long, about ⅛′ broad, the inner scales hirsute on the inner surface and along the margins and when fully grown often 1′ long, lustrous, brown above the middle, and reddish below. Bark ¼′—½′ thick, with a smooth light steel-gray surface. Wood hard, strong, tough, very close-grained, not durable, difficult to season, dark or often light red, with thin nearly white sapwood of 20—30 layers of annual growth; largely used in the manufacture of chairs, shoe-lasts, plane-stocks, the handles of tools, and for fuel. The sweet nuts are gathered and sold in the markets of Canada and of some of the western and middle states.

Distribution. Rich uplands and mountain slopes, often forming nearly pure forests, and southward on the bottom-lands of streams and the margins of swamps; valley of the Restigouche River, New Brunswick, to the northern shores of Lake Huron and the southern shores of Lake Superior, and southward to Virginia, Ohio, Michigan, the ravines of Rock River near Oregon, Ogle County, Illinois, Minnesota and northern Missouri; southward passing into the var. caroliniana Fern. & Rehd., differing in its ovate to short-ovate thicker leaves, usually rounded or subcordate at base, and often less coarsely serrate or undulate on the margins, glabrous or rarely densely soft pubescent below (f. mollis Fern. & Rehd.), in the often shorter involucre of the fruit with shorter and less crowded prickles; usually on the bottom-lands of streams and the borders of swamps, New Jersey, and southern Ohio and Missouri to western Florida, Alabama, Mississippi, Louisiana, eastern Texas, and northeastern Oklahoma; ascending on the southern Appalachian Mountains to altitudes of 3000°; probably growing to its largest size in eastern Louisiana.

The northern form is occasionally planted in the northern states as a shade and park tree.

2. Castanea Adans. Chestnut.

Trees or shrubs, with furrowed bark, porous brittle wood, durable in the ground, terete branchlets without terminal buds, axillary buds covered by 2 pairs of slightly imbricated scales, the outer lateral, the others accrescent, becoming oblong-ovate and acute and marking the base of the branch with narrow ring-like scars, and stout perpendicular tap-roots; producing when cut numerous stout shoots from the stump. Leaves convolute in the bud, ovate, acute, coarsely serrate, except at the base, with thin veins running to the points of the slender glandular teeth, deciduous; petioles leaving in falling small elevated semioval leaf-scars marked by an irregular marginal row of minute fibro-vascular bundle-scars; stipules ovate to linear-lanceolate, acute, scarious, infolding the leaf in the bud, caducous. Flowers opening in early summer, unisexual, strong-smelling; the staminate, in 3—7-flowered cymes, in the axils of minute ovate bracts, in elongated simple deciduous aments first appearing with the unfolding of the leaves from the inner scales of the terminal bud and from the axils of the lower leaves of the year, composed of a pale straw-colored slightly puberulous calyx deeply divided into 6 ovate rounded segments imbricated in the bud, and 10—20 stamens inserted on the slightly thickened torus, with filiform filaments incurved in the bud, becoming elongated and exserted, and ovoid or globose pale yellow anthers; the pistillate scattered or spicate at the base of the shorter persistent androgynous aments from the axils of later leaves, sessile, 2 or 3 together or solitary within a short-stemmed or sessile involucre of closely imbricated oblong acute bright green bracts scurfy-pubescent or tomentose below the middle, subtended by a bract and 2 lateral bractlets, each flower composed of an urn-shaped calyx, with a short limb divided into 6 obtuse lobes, minute sterile stamens shorter than the calyx-lobes, an ovary 6-celled after fecundation, with 6 linear spreading white styles hairy below the middle and tipped by minute acute stigmas, and 2 ovules in each cell, attached on its inner angle, descending, semianatropous. Fruit maturing in one season, its involucre inclosing 1—3 nuts, globose or short-oblong, pubescent or tomentose and spiny on the outer surface, with elongated ridged bright green ultimately brown branched spines fascicled between the deciduous scales, coated on the inner surface with lustrous pubescence, splitting at maturity into 2—4 valves; nut ovoid, acute, crowned by the remnants of the style, bright chestnut-brown and lustrous, tomentose or pubescent at apex, cylindrical, or when more than 1 flattened, marked at the broad base by a large conspicuous pale circular or oval thickened scar, its shell lined with rufous or hoary tomentum. Seed usually solitary by abortion, dark chestnut-brown, marked at apex by the abortive ovules, with thick and fleshy more or less undulate ruminate sweet farinaceous cotyledons.

Castanea is confined to the northern hemisphere, and is widely distributed through eastern North America, southern Europe, northern Africa, southwestern Asia, and central and northern China, Korea, and Japan. Seven species are distinguished. In the countries of the Mediterranean Basin much attention has been given to improving the fruit of the native species Castanea sativa Mill., which is occasionally planted in the middle United States; in Japan the seeds of Castanea crenata S. & Zucc. in many varieties and in China those of Castanea mollissima Bl. are important articles of food. Castanea produces coarse-grained wood very durable in contact with the soil, and rich in tannin. Chestnut-trees suffer in the eastern United States from the attacks of a fungus, Endothia parasitica Anders. which has nearly exterminated them in many parts of the country.

Castanea is the classical name of the Chestnut-tree.

Conspectus of the North American Species.

Involucre of the fruit containing 2 or 3 flattened nuts.

1. C. dentata (A, C).

Involucre of the fruit containing a single terete nut.

Involucre of the fruit densely covered with spines; branchlets hoary tomentose.

2. C. pumila (A, C).

Involucre of the fruit covered with scattered spines; branchlets glabrous or sparingly pilose.

3. C. alnifolia (C).

1. Castanea dentata Borkh. Chestnut.

Leaves oblong-lanceolate, acute and long-pointed at apex, gradually narrowed and cuneate at base, when they unfold puberulous on the upper surface and clothed on the lower with fine cobweb-like tomentum, at maturity thin, glabrous, dark dull yellow-green above, pale yellow-green below, 6′—8′ long, about 2′ wide, with a pale yellow midrib and primary veins; turning bright clear yellow late in the autumn; petioles stout, slightly angled, puberulous, ½′ long, often flushed with red; stipules ovate-lanceolate, acute, yellow-green, puberulous, about ½′ long. Flowers: staminate aments about ½′ long when they first appear, green below the middle and red above, becoming when fully grown 6′—8′ long, with stout green puberulous stems covered from base to apex with crowded flower-clusters; androgynous aments, slender, puberulous, 2½′—5′ long, with 2 or 3 irregularly scattered involucres of pistillate flowers near their base. Fruit: involucre attaining its full size by the middle of August, 2′—2½′ in diameter, sometimes a little longer than broad, somewhat flattened at apex, pubescent and covered on the outer surface with crowded fascicles of long slender glabrous much-branched spines, opening with the first frost and gradually shedding their nuts; nuts usually much compressed, ½′—1′ wide, usually rather broader than long, coated at apex or nearly to the middle with thick pale tomentum, the interior of the shell lined with thick rufous tomentum; seed very sweet.

A tree, occasionally 100° high, with a tall straight columnar trunk 3°—4° in diameter, or often when uncrowded by other trees with a short trunk occasionally 10°—12° in diameter, and usually divided not far above the ground into 3 or 4 stout horizontal limbs forming a broad low round-topped head of slightly pendulous branches frequently 100° across, and branchlets at first light yellow-green sometimes tinged with red, somewhat angled, lustrous, slightly puberulous, soon becoming glabrous and olive-green tinged with yellow or brown tinged with green and ultimately dark brown. Winter-buds ovoid, acute, about ¼′ long, with thin dark chestnut-brown scales scarious on the margins. Bark from 1′—2′ thick, dark brown and divided by shallow irregular often interrupted fissures into broad flat ridges separating on the surface into small thin closely appressed scales. Wood light, soft, not strong, liable to check and warp in drying, easily split, reddish brown, with thin lighter colored sapwood of 3 or 4 layers of annual growth; largely used in the manufacture of cheap furniture and in the interior finish of houses, for railway-ties, fence-posts, and rails. The nuts, which are superior to those of the Old World chestnuts in sweetness were formerly gathered in great quantities in the forest and sold in the markets of the eastern cities.

Distribution. Southern Maine to Woodstock, Grafton County, New Hampshire (rare) and to the valley of the Winooski River, Vermont, southern Ontario, and southern Michigan, southward to Delaware and Ohio, southern Indiana, and southwestern Illinois (Pulaski County) along the Appalachian Mountains up to altitudes of 4000° to northern Georgia, and to western Florida (Crestview, Walton County) southeastern (Henry and Dale Counties) and south central (Dallas County) Alabama, Northern, central and southeastern Mississippi (Pearl River County), and to central Kentucky and Tennessee; very common on the glacial drift of the northern states and, except at the north, mostly confined to the Appalachian hills; attaining its greatest size in western North Carolina and eastern Tennessee.

Formerly sometimes planted in the eastern states as an ornamental and timber tree, and for its nuts, of which several varieties have been recognized.

× Castanea neglecta Dode with leaves intermediate between those of C. dentata and C. pumila and an involucre containing a single large nut occurs on the Blue Ridge near Highlands, Macon County, North Carolina.

2. Castanea pumila Mill. Chinquapin.

Leaves oblong-elliptic to oblong-obovate, acute, coarsely serrate, with slender rigid spreading or incurved teeth, gradually narrowed and usually unequal and rounded or cuneate at base, when they unfold tinged with red and coated above with pale caducous tomentum and below with thick snowy white tomentum, at maturity rather thick and firm in texture, bright yellow-green on the upper surface, hoary or silvery pubescent on the lower, 3′—5′ long, 1½′—2′ wide; turning dull yellow in the autumn; petioles stout, pubescent, flattened on the upper side, ¼′—½′ long; stipules light yellow-green, pubescent, those of the 2 lowest leaves broad, ovate, acute, covered at apex by rufous tomentum, on later leaves ovate-lanceolate, often oblique and acute, becoming linear at the end of the branch. Flowers: staminate aments ½′ long when they first appear, pubescent, green below, bright red at apex, becoming when fully grown 4′—6′ long, with stout hoary tomentose stems and crowded or scattered flower-clusters; androgynous aments silvery tomentose, 3′—4′ long; involucres 1-flowered, scattered at the base of the ament or often spicate and covering its lower half, sessile or short-stalked. Fruit: involucre 1′—1½′ in diameter, with thin walls covered with crowded fascicles of slender spines tomentose toward the base; nut ovoid, terete, rounded at the slightly narrowed base, gradually narrowed and pointed at apex, more or less coated with silvery white pubescence, dark chestnut-brown, very lustrous, ¾′—1′ long, ⅓′ thick, with a thin shell lined with a coat of lustrous hoary tomentum, and a sweet seed.

A round-topped tree, rarely 50° high, with a short straight trunk 2°—3° in diameter, slender spreading branches, and branchlets coated at first with pale tomentum, becoming during their first winter pubescent or remaining tomentose at the apex, bright red-brown, glabrous, lustrous, olive-green or orange-brown during their second season and ultimately darker; east of the Mississippi River often a shrub spreading into broad thickets by prolific stolons, with numerous intricately branched stems often only 4° or 5° tall. Winter-buds ovoid, or oval, about ⅛′ long, clothed when they first appear in summer with thick hoary tomentum, becoming red during the winter and scurfy-pubescent. Bark ½′—1′ thick, light brown tinged with red, slightly furrowed and broken on the surface into loose plate-like scales. Wood light, hard, strong, coarse-grained, dark brown, with thin hardly distinguishable sapwood of 3 or 4 layers of annual growth; used for fence-posts, rails, and railway-ties. The sweet nuts are sold in the markets of the western and southern states.

Distribution. Dry sandy ridges, rich hillsides and the borders of swamps; southern New Jersey and Pennsylvania to central (Lake County) and western Florida and westward through the Gulf States to the valley of the Neches River, Texas, and through Arkansas to eastern Oklahoma and southwestern Missouri; on the Appalachian Mountains ascending to altitudes of 4500°; most abundant and of its largest size in southern Arkansas and eastern Texas.

3. Castanea alnifolia Nutt. Chinquapin.

A low shrub spreading into broad thickets by underground stems, with leaves pale pubescent on the lower surface; and distributed in the neighborhood of the coast from the valley of the Cape Fear River, North Carolina, to southern Georgia. Passing into

Castanea alnifolia var. floridana Sarg. Chinquapin.

Leaves oblong-obovate to elliptic, acute, acuminate or rounded at apex, gradually narrowed and cuneate or rounded at base, irregularly sinuate-toothed with apiculate teeth, hoary tomentose below when they unfold, soon glabrous with the exception of the last leaves of vigorous summer shoots, and at maturity thin, glabrous, dark green above, light green and lustrous below, 3′—4′ long and 1′—1¾′ wide; petioles stout, glabrous, about /12′ in length. Flowers: staminate aments pale pubescent, 4′—5′ long; androgynous aments pubescent, as long or rather longer with ten or twelve involucres of pistillate flowers below the middle, often only the lowest being fertilized. Fruit: involucre 1-seeded, subglobose to short-oblong, pale tomentose, ¾′ to 1¼′ in diameter, covered with stout pubescent scattered spines divided at base into numerous branches; nut ovoid, terete, acute, dark chestnut-brown, lustrous, ½′ to nearly ¾′ in length.

A tree occasionally 40°—45° high, with a tall trunk sometimes a foot in diameter, small irregularly spreading branches forming a narrow head, and slender glabrous or rarely pilose red-brown branchlets; more often a shrub sometimes with broader obovoid leaves sometimes puberulous on the lower surface.

Dry sandy soil; coast of North Carolina, near Wrightsville, New Hanover County; Dover, near the Ogechee River, Screven County, Georgia; Jacksonville, Duval County, and Panama City on Saint Andrew’s Bay, Bay County, Florida; near Selma, Dallas County, Alabama; and Covington, St. Tammany Parish, Louisiana.

A tree only on the shores of Saint Andrew’s Bay.

3. Castanopsis Spach.

Trees, with scaly bark, astringent wood, and winter-buds covered by numerous imbricated scales. Leaves convolute in the bud, 5-ranked, coriaceous, entire or dentate, penniveined, persistent; stipules obovate or lanceolate, scarious, mostly caducous. Flowers in 3-flowered cymes, or the pistillate rarely solitary or in pairs, in the axils of minute bracts, on slender erect aments from the axils of leaves of the year; the staminate on usually elongated and panicled aments, and composed of a campanulate 5 or 6-lobed or parted calyx, the lobes imbricated in the bud, usually 10 or 12 stamens inserted on the slightly thickened torus, with elongated exserted filiform filaments and oblong anthers, and a minute hirsute rudimentary ovary; the pistillate on shorter simple or panicled aments or scattered at the base of the staminate inflorescence, the cymes surrounded by an involucre of imbricated scales; calyx urn-shaped, the short limb divided into 6 obtuse lobes; abortive stamens inserted on the limb of the calyx and opposite its lobes; ovary sessile on the thin disk, 3-celled after fecundation, with 3 spreading styles terminating in minute stigmas, and 2 ovules in each cell attached to its interior angle. Fruit maturing at the end of the second or rarely of the first season, its involucre inclosing 1—3 nuts, ovoid or globose, sometimes more or less depressed, rarely obscurely angled, dehiscent or indehiscent, covered by stout spines, tuberculate or marked by interrupted vertical ridges; nut more or less angled by mutual pressure when more than 1, often pilose, crowned with the remnants of the style, marked at the base by a large conspicuous circular depressed scar, the thick shell tomentose on the inner surface. Seed usually solitary by abortion, bearing at apex the abortive ovules; cotyledons plano-convex, fleshy, farinaceous.

Castanopsis inhabits California with two species, and southeastern Asia where it is distributed with about twenty-five species from southern China to the Malay Archipelago and the eastern Himalayas. Of the California species one is usually arborescent and the other Castanopsis sempervirens Dudley is a low alpine shrub of the coast ranges and the Sierra Nevada.

Castanopsis from κὰστανα and ὄψις, in allusion to its resemblance to the Chestnut-tree.

1. Castanopsis chrysophylla A. DC. Chinquapin. Golden-leaved Chestnut.

Leaves lanceolate or oblong-ovate, gradually narrowed at the ends or sometimes abruptly contracted at apex into a short broad point, entire with slightly thickened revolute margins, when they unfold thin, coated below with golden yellow persistent scales and above with scattered white scales, at maturity thick and coriaceous, dark green and lustrous above, 2′—6′ long, ½′ to nearly 2′ wide, with a stout midrib raised and rounded on the upper side; turning yellow at maturity and falling gradually at the end of their second or in their third year; petioles ¼′—⅓′ in length; stipules ovate, rounded or acute at apex, brown and scarious, puberulous, ¼′—⅓′ long. Flowers appearing irregularly from June until February in the axils of broadly ovate apiculate pubescent bracts on staminate and androgynous scurfy stout-stemmed aments 2′—2½′ long and crowded at the ends of the branches; calyx of the staminate flower coated on the outer surface with hoary tomentum, divided into broadly ovate rounded lobes much shorter than the slender stamens; calyx of the pistillate flower oblong-campanulate, free from the ovary, clothed with hoary tomentum, divided at apex into short rounded lobes, rather shorter than the minute abortive stamens; anthers red; ovary conic, hirsute, with elongated slightly spreading thick pale stigmas. Fruit ripening at the end of the second season, involucre globose, dehiscent, irregularly 4-valved, often slightly shorter than the nuts, sessile, solitary, or clustered, tomentose and covered on the outer surface by long stout or slender rigid spines, 1′—1½′ in diameter, containing 1 or occasionally 2 nuts; nuts broadly ovoid, acute, obtusely 3-angled, light yellow-brown and lustrous; seeds dark purple-red, sweet and edible.

A tree, 50°—100° high, with a massive trunk 3°—6° in diameter, frequently free of branches for 50°, stout spreading branches forming a broad compact round-topped or conic head, and rigid branchlets coated when they first appear with bright golden-yellow scurfy scales, dark reddish brown and slightly scurfy during their first winter, and gradually growing darker in their second season; often much smaller and sometimes reduced to a shrub, 2°—12° high (var. minor A. De Candolle). Winter-buds fully grown at mid-summer, usually crowded near the end of the branch, ovoid or subglobose, with broadly ovate apiculate thin and papery light brown scales slightly puberulous on the back, ciliate on the scarious often reflexed margins, the terminal about ¼′ long and broad and rather larger than the often stipitate axillary buds. Bark 1′—2′ thick and deeply divided into rounded ridges 2′—3′ wide, broken into thick plate-like scales, dark red-brown on the surface and bright red internally. Wood light, soft, close-grained, not strong, light brown tinged with red, with thin lighter colored sapwood of 50—60 layers of annual growth; occasionally used in the manufacture of ploughs and other agricultural implements.

Distribution. Skamania County, Washington, valley of the lower Columbia River, Oregon, southward along the western slopes of the Cascade Mountains, and in California along the western slopes of the Sierra Nevada and through the coast ranges to the elevated valleys of the San Jacinto Mountains, sometimes ascending to altitudes of 4000° above the sea; of its largest size in the humid coast valleys of northern California.

Occasionally cultivated in the gardens of temperate Europe.

4. Lithocarpus Bl.

Pasania Örst.

Trees, with astringent properties, pubescence of fascicled hairs, deeply furrowed scaly bark, hard close-grained brittle wood, stout branchlets, and winter-buds covered by few erect or spreading foliaceous scales. Leaves convolute in the bud, petiolate, persistent, entire or dentate, with a stout midrib, primary veins running obliquely to the points of the teeth, or on entire leaves forked and united near the margins, and reticulate veinlets; stipules oblong-obovate to linear-lanceolate, those of the upper leaves persistent and surrounding the buds during the winter. Flowers in erect unisexual and in bisexual tomentose aments from the axils of leaves of the year, from the inner scales of the terminal bud or from separate buds in the axils of leaves of the previous year; staminate in 3-flowered clusters in the axils of ovate rounded bracts, the lateral flowers subtended by similar but smaller bracts, each flower composed of a 5-lobed tomentose calyx, with nearly triangular acute lobes, 10 stamens, with slender elongated filaments and small oblong or emarginate anthers, and an acute abortive hairy ovary; pistillate scattered at the base of the upper aments below the staminate flowers, solitary in the axils of acute bracts, furnished with minute lateral bractlets, and composed of a 6-lobed ovoid calyx, with rounded lobes, inclosed in the tomentose involucral scales, 6 stamens, with abortive anthers, an ovoid-oblong 3-celled ovary, 3 elongated spreading light green styles thickened and stigmatic at apex, and 2 anatropous ovules in each cell. Fruit an oval or ovoid nut maturing at the end of the second season, 1-seeded by abortion, surrounded at base by the accrescent woody cupular involucre of the flower, marked by a large pale circular basal scar, the thick shell tomentose on the inner surface. Seed red-brown, filling the cavity of the nut, bearing at apex the abortive ovules; cotyledons thick and fleshy, yellow and bitter.

Lithocarpus is intermediate between the Oaks and the Chestnuts, and, with the exception of one California species, is confined to southeastern Asia, where it is distributed with many species from southern Japan and southern China through the Malay Peninsula to the Indian Archipelago.

Lithocarpus from λίθος and καρπός, in allusion to the character of the fruit.

1. Lithocarpus densiflora Rehd. Tan Bark Oak. Chestnut Oak.

Quercus densiflora Hook. & Arn.
Pasania densiflora Örst.

Leaves oblong or oblong-obovate, rounded or acute or rarely cordate at base, acute or occasionally rounded at apex, or rarely lanceolate and acuminate (f. lanceolata Rehdr.) repand-dentate, with acute callous teeth, or entire with thickened revolute margins, coated when they unfold with fulvous tomentum and glandular on the margins with dark caducous glands, at maturity pale green, lustrous and glabrous or covered with scattered pubescence on the upper surface, rusty-tomentose on the lower, ultimately becoming glabrous above and glabrate and bluish white below, 3′—5′ long, ¾′—3′ wide, with a midrib raised and rounded on the upper side, thin or thick primary veins and fine conspicuous reticulate veinlets; persistent until the end of their third or fourth year; petioles stout, rigid, tomentose, ½′—¾′ in length; stipules brown and scarious, hirsute on the outer surface. Flowers in early spring and frequently also irregularly during the autumn; aments stout-stemmed, 3′—4′ long; staminate flowers crowded, hoary-tomentose in the bud, their bracts tomentose. Fruit solitary or often in pairs, on a stout tomentose peduncle ½′—1′ in length; nut full and rounded at base, gradually narrowed and acute or rounded at apex, scurfy-pubescent when fully grown, becoming light yellow-brown, glabrous and lustrous at maturity, ¾′—1′ long, ½′—1′ thick, its cup shallow, tomentose with lustrous red-brown hairs on the inner surface, and covered by long linear rigid spreading or recurved light brown scales coated with fascicled hairs, frequently tipped, especially while young, with dark red glands and often tomentose near the base of the cup.

A tree, usually 70°—80° but sometimes 150° high, with a trunk 1°—4° in diameter, stout branches ascending in the forest and forming a narrow spire-like head, or in open positions spreading horizontally and forming a broad dense symmetrical round-topped crown, and branchlets coated at first with a thick fulvous tomentum of fascicled hairs often persistent until the second or third year, becoming dark reddish brown and frequently covered with a glaucous bloom; or sometimes reduced to a shrub, with slender stems only a few feet high (var. montana Rehdr.). Winter-buds ovoid, obtuse, ¼′—⅓′ long, often surrounded by the persistent stipules of the upper leaves, with tomentose loosely imbricated scales, those of the outer ranks linear-lanceolate, increasing in width toward the interior of the bud, those of the inner ranks ovate or obovate and rounded at apex. Bark ¾′—1½′ thick, deeply divided by narrow fissures into broad rounded ridges broken into nearly square plates covered by closely appressed light red-brown scales. Wood hard, strong, close-grained, brittle, reddish brown, with thick darker brown sapwood; largely used as fuel. The bark is exceedingly rich in tannin and is largely used for tanning leather.

Distribution. Valley of the Umpqua River, Oregon, southward through the coast ranges to the Santa Inez Mountains, California, and along the western slope of the Sierra Nevada up to elevations of 4000° above the sea to Mariposa County; very abundant in the humid coast region north of San Francisco Bay and on the Santa Cruz and Santa Lucia Mountains, and of its largest size in the Redwood forest of Napa and Mendocino Counties; southward and on the Sierras less abundant and of smaller size; the form lanceolata in southern Oregon and in Del Norte and Mendocino Counties, California; the var. montana at high altitudes on the Siskiyou Mountains, in the region of Mount Shasta and on the northern Sierra Nevada.

5. Quercus L. Oak.

Trees or shrubs, with astringent properties, pubescence of fascicled hairs, scaly or dark and furrowed bark, hard and close-grained or porous brittle wood, slender branchlets marked by pale lenticels and more or less prominently 5-angled. Winter-buds clustered at the ends of the branchlets, with numerous membranaceous chestnut-brown slightly accrescent caducous scales closely imbricated in 5 ranks, in falling marking the base of the branchlet with ring-like scars. Leaves 5-ranked, lobed, dentate or entire, often variable on the same branch, membranaceous or coriaceous, the primary veins prominent and extending to the margins or united within them and connected by more or less reticulate veinlets, deciduous in the autumn or persistent until spring or until their third or fourth year; petioles in falling leaving slightly elevated semiorbicular more or less obcordate leaf-scars broader than high, marked by the ends of numerous scattered fibro-vascular bundles; stipules obovate to lanceolate, scarious, caducous, or those of upper leaves occasionally persistent through the season. Flowers vernal with or after the unfolding of the leaves; staminate solitary in the axils of lanceolate acute caducous bracts, or without bracts, in graceful pendulous clustered aments, from separate or leaf-buds in the axils of leaves of the previous year, or from the axils of the inner scales of the terminal bud or from those of the leaves of the year; calyx campanulate, lobed or divided to the base into 4—7, usually 6, membranaceous lobes; stamens 4—6, rarely 2, or 10—12, inserted on the slightly thickened torus, with free filiform exserted filaments and ovate-oblong or subglobose glabrous or rarely hairy 2-celled usually yellow anthers; pistillate solitary, subtended by a caducous bract and 2 bractlets, in short or elongated few-flowered spikes from the axils of leaves of the year; calyx urn-shaped, with a short campanulate 6-lobed limb, the tube adnate to the incompletely 3 or rarely 4 or 5-celled ovary inclosed more or less completely by an accrescent involucre of imbricated scales, becoming the cup of the fruit; styles as many as the cells of the ovary, short or elongated, erect or incurved, dilated above, stigmatic on the inner face or at apex only, generally persistent on the fruit; ovules anatropous or semianatropous, 2 in each cell. Fruit a nut (acorn) maturing in one or in two years, ovoid, subglobose, or turbinate, short-pointed at apex, 1-seeded by abortion, marked at base by a large conspicuous circular scar, with a thick shell, glabrous or coated on the inner surface with pale tomentum, more or less surrounded or inclosed in the accrescent cupular involucre of the flower (cup), its scales thin or thickened, loosely or closely imbricated. Seed marked at base or at apex or rarely on the side by the abortive ovules; cotyledons thick and fleshy, usually plano-convex and entire.

Quercus inhabits the temperate regions of the northern hemisphere and high altitudes within the tropics, ranging in the New World southward to the mountains of Colombia and in the Old World to the Indian Archipelago. Two hundred and seventy-five species have been described; of the North American species fifty-four are large or small trees. Of exotic species, the European Quercus Robur L., and Quercus sessiliflora Salisb., have been frequently cultivated as ornamental trees in the eastern United States, where, however, they are usually short-lived and unsatisfactory. Many of the species are important timber-trees; their bark is often rich in tannin and is used for tanning leather, and all produce wood valuable for fuel and in the manufacture of charcoal.

Quercus is the classical name of the Oak-tree.

Conspectus of the North American Arborescent Species.

Fruit maturing at the end of the second season (except 22); shell of the nut silky tomentose on the inner surface; leaves or their lobes bristle-tipped. Black Oaks.

Stamens usually 4—6; styles elongated, finally recurved; abortive ovules apical.

Leaves deciduous in their first autumn or winter.

Leaves pinnately lobed, convolute in the bud.

Leaves green on both surfaces.

Scales of the cup of the fruit closely appressed.

Leaves usually dull on the upper surface, 7—11-lobed; cup of the fruit cup-shaped or in one variety broad and saucer-shaped, its scales thin.

1. Q. borealis (A).

Leaves lustrous.

Leaves dimorphous, 5—7-lobed, axillary clusters of hairs large and prominent; cup of the fruit saucer-shaped or in one form deep cup-shaped.

2. Q. Shumardii (A, C).

Leaves similar on upper and lower branches.

Cup of the fruit turbinate or deep cup-shaped.

Leaves 5-lobed, the lobes usually entire, rarely furnished with tufts of axillary hairs below.

3. Q. texana (C).

Leaves 5—7-lobed, the lobes dentate, furnished with tufts of axillary hairs below.

4. Q. ellipsoidalis (A).

Cup of the fruit deep cup-shaped to turbinate; leaves 5—9-lobed, the lobes toothed.

5. Q. coccinea (A, C).

Cup of the fruit saucer-shaped.

Leaves 5—9-lobed.

6. Q. palustris (A, C).

Leaves 3—5-lobed.

7. Q. georgiana (C).

Scales of the cup of the fruit more or less loosely imbricated, forming a free margin on its rim.

Leaves usually 7-lobed.

Winter-buds tomentose.

8. Q. velutina (A, C).

Winter-buds pubescent only at apex.

9. Q. Kelloggii (G).

Leaves usually 3—5-lobed; winter-buds rusty pubescent.

10. Q. Catesbæi (C).

Leaves whitish or grayish tomentulose below.

Leaves mostly acutely 5-lobed, pale or silvery white below.

11. Q. ilicifolia (A).

Leaves often dimorphous, 3—11-lobed, the lobes often falcate.

12. Q. rubra (A, C).

Leaves broad-obovate, often abruptly dilated at the wide obscurely lobed apex.

Leaves rounded or cordate at base.

Lower surface of the leaves orange color or brownish, the upper scales of the cup forming with several rows a thick rim on its inner surface, often reflexed.

13. Q. marilandica (A, C).

Lower surface of the leaves pale, the erect scales on the rim of the cup in a single row.

14. Q. arkansana (C).

Leaves cuneate at base.

Leaves oblong-obovate.

15. Q. nigra (C).

Leaves rhombic.

16. Q. rhombica (C).

Leaves lanceolate-oblong or lanceolate-obovate, usually entire, involute in the bud. Willow Oaks.

Leaves glabrous.

Leaves lanceolate to oblanceolate, deciduous in autumn.

17. Q. Phellos (A, C).

Leaves elliptic or rarely oblong-obovate, deciduous in the late winter.

18. Q. laurifolia (C).

Leaves tomentose or pubescent below, oblong-lanceolate to oblong-obovate.

Leaves pale blue-green, hoary tomentose below.

19. Q. cinerea (C).

Leaves dark green, pubescent below.

20. Q. imbricaria (A).

Leaves not deciduous in the autumn, revolute in the bud (convolute in 23).

Leaves mostly persistent until after the appearance of those of the following year.

Leaves lanceolate, oblong-lanceolate or elliptic, pale and tomentose below.

21. Q. hypoleuca (E, H).

Leaves oval, orbicular to oblong, green and pubescent below; fruit maturing at the end of the first season.

22. Q. agrifolia (G).

Leaves persistent until their second summer or autumn.

Leaves lanceolate to oval or oblong-lanceolate, entire or serrate; cup of the fruit turbinate or tubular.

23. Q. Wislizenii (G).

Leaves oval to oblong-obovate; cup of the fruit saucer-shaped or turbinate.

24. Q. myrtifolia (C).

Stamens usually 6—8; styles dilated; abortive ovules basal or lateral; leaves persistent until their third or fourth season, involute in the bud.

Leaves oblong, entire, dentate, or sinuate-toothed, fulvous-tomentose and ultimately pale on the lower surface; cup of the fruit usually thick.

25. Q. chrysolepis (G, H).

Leaves oblong-lanceolate, crenate-dentate or entire, pubescent or tomentose below; cup of the fruit usually thin.

26. Q. tomentella (G).

Fruit maturing at the end of the first season; shell of the nut glabrous on the inner surface (hoary-tomentose in 27); abortive ovules basal; stamens 6—8; styles dilated; lobes of the leaves not bristle-tipped. White Oaks.

Leaves mostly persistent until the appearance of those of the following year, revolute in the bud (convolute in 28).

Leaves yellow-green.

Fruit sessile or short-stalked.

Leaves oblong-lanceolate, entire or repand-dentate; inner surface of the nut hoary tomentose.

27. Q. Emoryi (F, H).

Leaves oblong or obovate, entire, sinuate-toothed or lobed.

28. Q. dumosa (G).

Fruit long-stalked; leaves oblong, elliptic or obovate, pale, glabrous or in one form densely tomentose below.

29. Q. virginiana (C).

Leaves blue-green.

Fruit usually in many-fruited long-stalked clusters; leaves broad-obovate, coarsely reticulate-venulose.

30. Q. reticulata (H).

Fruit solitary or in pairs.

Cup of the fruit saucer-shaped; leaves ovate to ovate-oblong, entire.

31. Q. Toumeyi (H).

Cup of the fruit cup-shaped or hemispherical, oblong-lanceolate to broad-obovate, pubescent below.

32. Q. arizonica (H).

Cup of the fruit usually cup-shaped or turbinate.

Leaves ovate, oval or obovate, usually cordate at base; fruit rather long-stalked.

33. Q. oblongifolia (E, H).

Leaves oblong to obovate, usually cuneate or rounded or cordate at base.

34. Q. Engelmannii (G).

Leaves deciduous in their first season.

Leaves blue-green.

Arboreous; leaves oblong, lobed, spinescent-dentate or entire, pubescent below; cup of the fruit shallow cup-shaped.

35. Q. Douglasii (G).

Arborescent or shrubby.

Leaves oblong to oblong-obovate, undulate-lobed; cup of the fruit saucer-shaped to cup-shaped.

36. Q. Vaseyana (C).

Leaves oblong-obovate to elliptic or lanceolate, undulate, serrate-toothed or irregularly lobed; cup of the fruit hemispheric to cup-shaped.

37. Q. Mohriana (C).

Leaves oblong to oblong-ovate, slightly lobed or entire; cup of the fruit cup-shaped or rarely saucer-shaped.

38. Q. Laceyi (C).

Leaves yellow-green.

Leaves entire or slightly lobed.

Leaves different on upper and lower branches, oblong to oblong-obovate, slightly lobed or entire.

Cup of the fruit cup-shaped.

39. Q. annulata (C).

Cup of the fruit shallow saucer-shaped.

40. Q. Durandii (C).

Leaves similar on upper and lower branches, entire or slightly sinuate-lobed toward the apex, oblong or oblong-obovate.

41. Q. Chapmanii (C).

Leaves more or less deeply sinuate-lobed.

Leaves white-tomentulose below (sometimes green and pubescent in 43).

Leaves obovate or oblong, lyrately pinnatifid or deeply sinuate-lobed; cup of the fruit fringed by the awned scales.

42. Q. macrocarpa (A, C, F).

Leaves obovate-oblong, deeply 5—9-lobed or pinnatifid; nut often inclosed in the cup.

43. Q. lyrata (A, C).

Leaves pubescent below.

Leaves usually covered above with fascicled hairs, obovate, 3—5-lobed, their lobes truncate or rounded.

44. Q. stellata (A, C).

Leaves glabrous above at maturity.

Leaves obovate to oblong; cup of the fruit shallow cup-shaped or slightly turbinate, its scales usually thin.

45. Q. Garryana (B, G).

Leaves oblong-obovate; cup of the fruit hemispheric, the scales often much thickened.

46. Q. utahensis (F).

Leaves oblong-obovate, deeply lobed; nut conic, elongated, inclosed for one-third its length in the cup-shaped cup.

47. Q. lobata (G).

Leaves glabrate or puberulous below, oblong to oblong-obovate.

48. Q. leptophylla (F).

Leaves glabrous below.

Leaves oblong-obovate, usually 5-lobed.

49. Q. austrina (C).

Leaves oblong-obovate, obliquely pinnatifid or 3—9-lobed.

50. Q. alba (A, C).

Leaves coarsely sinuate-toothed. Chestnut Oaks.

Fruit on peduncles much longer than the petioles; leaves obovate or oblong-obovate, generally sinuate-dentate or lobed, pubescent, and usually hoary on the lower surface.

51. Q. bicolor (C).

Fruit on peduncles about as long or shorter than the petioles.

Leaves obovate or oblong-obovate, cuneate or rounded at the broad or narrow base, tomentose or pubescent and often silvery white below.

52. Q. Prinus (A, C).

Leaves obovate or oblong to lanceolate, acuminate, with rounded or acute teeth.

53. Q. montana (A, C).

Fruit sessile or nearly so; leaves oblong to lanceolate, acute or acuminate or broadly obovate, puberulous and pale, often silvery white on the lower surface.

54. Q. Muehlenbergii (A, C).

1. Quercus borealis Michx. Red Oak.

Leaves obovate or oblong, acute or acuminate, abruptly or gradually cuneate or rounded at the broad or narrow base, usually divided about half way to the midrib by wide oblique sinuses rounded at the bottom into 11 or sometimes into 7 or 9 acute oblique ovate lobes tapering from broad bases and mostly sinuately 3-toothed at apex with elongated bristle-pointed teeth, or sometimes oblong-obovate, gradually narrowed and cuneate at base, and sinuately lobed with broad acute usually entire or slightly dentate lobes, when they unfold pink, covered with soft silky pale pubescence on the upper surface and below with thick white tomentum, soon glabrous, at maturity thin and firm, dark green, dull and glabrous above, pale yellow-green, glabrous or rarely puberulous and sometimes furnished with small tufts of rusty hairs in the axils of the veins below, 5′—9′ long, 4′—6′ wide; falling early in the autumn after turning dull or sometimes bright orange color or brown; petioles stout, yellow or red, 1′—2′ in length. Flowers: staminate in pubescent aments 4′—5′ long; calyx divided into 4 or 5 narrow ovate rounded lobes shorter than the stamens; pistillate on short glabrous peduncles, their involucral scales broadly ovate, dark reddish brown, shorter than the conspicuous linear acute bract of the flower and as long as the lanceolate acute calyx-lobes; stigmas bright green. Fruit solitary or in pairs, sessile or short-stalked, ovoid, gradually narrowed and acute at apex or cylindric and rounded at apex, pale brown, lustrous, more or less tomentose toward the ends, ½′—1′ long; ½′—¾′ in diameter; cup cup-shaped, puberulous on the inner surface, covered with small closely appressed ovate acute red-brown pubescent scales slightly thickened on the back toward the base of the cup, with a thin dark-colored tip and margins.

A tree usually not more than 60°—70° high, with a trunk 2°—3° in diameter, often much smaller, stout branches forming a narrow head, and slender lustrous branchlets light green and covered with pale scurfy pubescence when they first appear, dark red during their first winter and ultimately dark brown. Winter-buds ovoid, gradually narrowed to the acute apex, about ¼′ long, with thin ovate acute light chestnut-brown scales. Bark on young stems and on the upper part of the limbs of old trees 1′—1½′ thick, dark brown tinged with red and divided into small thick appressed plates scaly on the surface. Wood heavy, hard, strong, close-grained, light reddish brown, with thin lighter-colored sapwood; used in construction, for the interior finish of houses, and in furniture.

Distribution. Nova Scotia and New Brunswick, through Quebec to southern Ontario, and southward to northern New England, western New York, northern Pennsylvania (Presque Isle, Erie County), northern Michigan, southeastern Wisconsin, central Minnesota, central Iowa (Winneshick County), and on the Appalachian Mountains of North Carolina at altitudes of about 4000°. Passing with many intermediate forms differing in the size of the nut and in the depth of the cup into

Quercus borealis var. maxima Ashe. Red Oak.

Quercus rubra Du Roi, not L.

Fruit solitary or in pairs, sessile or short-stalked; nut ovoid to slightly obovoid, gradually narrowed and rounded at apex, slightly narrowed at base, usually 1′—1¼′ long and ½′—⅔′ thick, occasionally not more than ⅔′ long and thick, inclosed only at the base in a thick saucer-shaped cup.

A tree, usually 70°—80°, or occasionally 150° high, with a trunk 3°—4° in diameter, and stout spreading and ascending branches forming a broad head.

Distribution. Province of Quebec in the neighborhood of Montreal, and southern Ontario, westward through southern Michigan to southeastern Nebraska, and southward to northern Georgia, on the southern Appalachian Mountains up to altitudes of 3000°, southern Kentucky, eastern and central Tennessee, northeastern (Tishomingo County), northwestern (Yazoo County), and central and southern (Hinds and Union Counties) Mississippi, northern and southwestern Alabama (Dekalb, Cullman, Jefferson, and Dallas Counties), northwestern Arkansas, and eastern Kansas and Oklahoma; one of the largest and most generally distributed trees of the northern states; rare and local in the south; of its largest size in the region north of the Potomac and Ohio Rivers.

Often planted as a park and shade tree in the northeastern states and in the counties of western and northern Europe; generally more successful in Europe than other American Oaks.

× Quercus Lowellii Sarg., a possible hybrid of Quercus borealis and Q. ilicifolia, has been found in the neighborhood of Seabury, York County, Maine.

× Quercus Porterii Trel., probably a hybrid of Quercus borealis var. maxima and Q. velutina, has been found on Bowditch Hill, Jamaica Plain, Suffolk County, Massachusetts, on College Hill, Easton, Northampton County, Pennsylvania, and near Columbus, Franklin County, Ohio.

× Quercus runcinata Engelm., believed to be a hybrid of Quercus borealis var. maxima and Q. imbricaria first found near St. Louis, occurs also in the neighborhood of Independence, Jackson County, and at Williamsville, Wayne County, Missouri, and in Richland and Wayne Counties, Illinois.

2. Quercus Shumardii Buckl.

Quercus texana Sarg. in part, not Buckl.

Leaves obovate, seven rarely five-lobed, the lobes two or three-lobed and sometimes dentate at apex, on leaves of lower branches short and broad, and separated by narrow sinuses pointed or rounded in the bottom, on upper branches deeply divided by broad rounded sinuses into narrow acuminate lobes, when they unfold often tinged with red and covered with pale loose tomentum deciduous before they are half grown, at maturity glabrous, dark green and lustrous above, paler and furnished below with large axillary tufts of pale hairs, 6′—8′ long, 4′—5′ wide, with a thin midrib and slender primary veins running to the points of the lobes; petioles slender, glabrous, 2′—2½′ in length. Flowers: staminate in slender glabrous aments 6′—7′ long; calyx divided into 4 or 5 rounded slightly villose lobes shorter than the stamens; pistillate on pubescent peduncles, their involucral scales ovate, light brown, pubescent; stigmas red. Fruit: nut oblong-ovoid, narrowed and rounded at apex, ¾′—1¼′ long, ½′—1′ in diameter, inclosed at the base only in the thick saucer-shaped cup with a slightly incurved rim and covered with closely appressed ovate pale pubescent or nearly glabrous scales narrowed above the middle, abruptly long-pointed, thin or often conspicuously tuberculate.

A tree up to 120° high, with a tall trunk occasionally 5° in diameter, stout wide-spreading branches forming a broad rather open head, and gray or grayish brown glabrous branchlets. Winter-buds ovoid, acute or acuminate, about ¼′ long, with closely imbricated gray glabrous or rarely pubescent scales. Bark 1′—1½′ thick, ridged, broken into small appressed plates scaly on the surface. Wood heavy, hard, close-grained, light reddish brown, often manufactured into lumber in the Mississippi valley and considered more valuable than that of the northern Red Oak.

Distribution. Borders of streams and swamps in moist rich soil; coast region of Texas eastward from the Colorado River and ranging inland up the valley of that river to Burnet County, southeastern Oklahoma, through Arkansas, southeastern Kansas and Missouri to Fayette County, Iowa, southern Illinois and Indiana, the neighborhood of Columbus, Franklin County, Ohio, and southeastern Michigan (near Portage Lake, Jackson County); through the eastern Gulf States to western and central Florida and northward in the neighborhood of the coast to the valley of the Neuse River, North Carolina; Chesapeake Beach, Calvert County, Maryland (W. W. Ashe); ranging inland in the south Atlantic States to Rome, Floyd County, Georgia, Calhoun Falls, Abbeville County, and Columbia, Richland County, South Carolina, and Chapel Hill, Orange County, North Carolina. Passing into

Quercus Shumardii var. Schneckii Sarg.

Quercus texana Sarg. in part, not Buckl.
Quercus Schneckii Britt.

Differing from the type in the deep cup-shaped cup of the fruit covered with thin scales, rarely much thickened and tuberculate at base (only on river banks near Vicksburg, Warren County, Mississippi), and connected with it by forms with the cups of the fruit differing from saucer to deep cup-shaped.

Distribution. Growing with Quercus Shumardii; more common in Texas and in the Mississippi valley than the type, and ranging eastward through Louisiana and Mississippi to central and southern Alabama, central and southeastern Tennessee (neighborhood of Chattanooga), and central Kentucky; apparently not reaching the Atlantic States.

3. Quercus texana Buckl.

Leaves widest above the middle, broad-cuneate, concave-cuneate or nearly truncate at base, deeply or rarely only slightly divided by broad sinuses rounded in the bottom into 5 or 7 lobes, the terminal lobe 3-lobed and acute at apex, the upper lateral lobes broad and more or less divided at apex and much larger and more deeply lobed than those of the lowest pair, when they unfold densely covered with fascicled hairs and often bright red, soon glabrous, thin, dark green and lustrous above, pale and lustrous and rarely furnished below with small inconspicuous axillary tufts of pale hairs, 3′—3½′ long, 2½′—3′ wide, with a thin midrib and slender primary veins running to the points of the lobes; petioles slender, soon glabrous, ¾′—1½′ in length. Flowers: staminate in slender villose aments 3′—4′ long; calyx thin, villose on the outer surface, divided into 4 or 5 acute lobes shorter than the stamens; pistillate on short hoary tomentose peduncles, their involucral scales brown tinged with red; stigmas bright red. Fruit short-stalked, usually solitary; nut ovoid, narrowed and rounded at apex, light red-brown, often striate, ¼′—¾′ long and broad, sometimes acute, nearly 1′ in length and not more than ⅓′ in diameter; cup turbinate, covered with thin ovate acuminate slightly appressed glabrous scales, in the small fruit of trees on dry hills inclosing a third or more of the nut, in the larger fruit of trees on better soil comparatively less deep.

A tree on dry hills rarely more than 30° tall, with a trunk 8′—10′ in diameter, small spreading or erect branches and slender red or reddish brown glabrous or rarely pubescent branchlets; often a shrub; on better soil at the foot of hills occasionally 50° high with a trunk 12′—18′ in diameter. Winter-buds ovoid, acute, ⅙′—¼′ long and covered with closely imbricated acute slightly or densely pubescent red scales. Bark light brown tinged with red, ¾′—1′ thick, deeply ridged and broken into plate-like scales.

Distribution. Dry limestone hills and ridges, and in the more fertile soil at their base; central and western Texas (Dallas, Tarrant County to Travis and Bexar Counties), and to the Edwards Plateau (San Saba, Kerr, Brown, Coke and Uvalde Counties); westward replaced by the var. chesosensis Sarg. differing in the acuminate lobes of the leaves and smaller cups of the fruit; known only on the dry rocky slopes of the Chesos Mountains, Brewster County, Texas; and by the var. stellapila Sarg., differing in the presence of fascicled hairs on both surfaces of the mature leaves and on the branchlets of the year; above Fort Davis, Jeff Davis County, Texas.

4. Quercus ellipsoidalis E. J. Hill. Black Oak.

Leaves elliptic to obovate-orbicular, acute or acuminate, truncate or broadly cuneate at base, deeply divided by wide sinuses rounded in the bottom into 5—7 oblong lobes repandly dentate at apex, or often, especially those of the upper pair, repandly lobulate, when they unfold slightly tinged with red and hoary-tomentose, soon becoming glabrous with the exception of small tufts of pale hairs in the axils of the principal veins, at maturity thin and firm, bright green and lustrous above, paler and sometimes entirely glabrous below, 3′—5′ long, 2½′—4′ wide, with a stout midrib and primary veins and prominent reticulate veinlets; late in the autumn turning yellow or pale brown more or less blotched with purple; petioles slender, glabrous or rarely puberulous, 1½′—2′ in length. Flowers: staminate in puberulous aments 1½′—2′ long; calyx campanulate, usually tinged with red, 2—5-lobed or parted into oblong-ovate or rounded segments, glabrous or slightly villose, fringed at apex with long twisted hairs, about as long as the 2—5 stamens, with short filaments and oblong anthers; pistillate on stout tomentose 1—3-flowered peduncles, red, their involucral scales broad, oblong, acute, hairy; calyx campanulate, 4—7-lobed, ciliate on the margins. Fruit short-stalked or nearly sessile, solitary or in pairs; nut ellipsoidal to subglobose, chestnut-brown, often striate and puberulous, inclosed for one third to one half its length in a turbinate or cup-shaped cup gradually narrowed at base, thin, light red-brown, and covered by narrow ovate obtuse or truncate brown pubescent closely appressed scales.

A tree, 60°—70° high, with a short trunk rarely 3° in diameter, much forked branches ascending above and often pendulous low on the stem, forming a narrow oblong head, and slender branchlets covered at first with matted pale hairs, bright reddish brown during their first winter, becoming dark gray-brown or reddish brown in their second season. Winter-buds ovoid, obtuse or acute, sometimes slightly angled, about ⅛′ long, with ovate or oval red-brown lustrous slightly puberulous outer scales ciliate on the margins. Bark thin, light yellow internally, close, rather smooth, divided by shallow connected fissures into thin plates, dark brown near the base of the tree, dull above, gray-brown and only slightly furrowed on the large branches.

Distribution. In the neighborhood of Chicago, Illinois, to southeastern Minnesota common; often covering large areas of sandy soil with a stunted growth and on the prairies sometimes a low shrub; eastern Iowa (Muscatine County), and the Lower Peninsular of Michigan (Montmorency, Arenac, and St. Clair Counties).

5. Quercus coccinea Muench. Scarlet Oak. Spanish Oak.

Leaves oblong-obovate or elliptic, truncate or cuneate at base, deeply divided by wide sinuses rounded in the bottom into 7 or rarely 9 lobes repand-dentate at apex, the terminal lobe, ovate, acute, and 3-toothed, the middle division the largest and furnished with 2 small lateral teeth, the lateral lobes obovate, oblique or spreading, sometimes falcate, usually broad and oblique at the coarsely toothed apex, when they unfold bright red covered with loose pale pubescence above and below with silvery white tomentum, green at the end of a few days, at maturity thin and firm, bright green, glabrous and very lustrous above, paler and less lustrous and sometimes furnished with small tufts of rusty pubescence in the axils of the veins below, 3′—6′ long, 2½′—4′ broad, with a yellow midrib and primary veins, late in the autumn turning brilliant scarlet; petioles slender, terete, 1½′—2½′ in length. Flowers: staminate in slender glabrous aments 3′—4′ long; calyx pubescent, bright red before opening, divided into 4 or 5 ovate acute segments shorter than the stamens; pistillate on pubescent peduncles sometimes ½′ long, bright red, their involucral scales ovate, pubescent, shorter than the acute calyx-lobes. Fruit sessile or stalked, solitary or in pairs; nut oval, oblong-ovate or hemispheric, truncate or rounded at base, rounded at apex, ½′—1′ long, ⅓′—⅔′ thick, light reddish brown and occasionally striate, inclosed for one third to one half its length in a deep cup-shaped or turbinate thin cup light reddish brown on the inner surface, covered by closely imbricated oblong-ovate acute thin, or rarely much thickened (var. tuberculata Sarg.) light reddish brown slightly puberulous scales.

A tree, 70°—80° high, with a trunk 2°—3° in diameter, comparatively small branches spreading gradually and forming a rather narrow open head, and slender branchlets coated at first with loose scurfy pubescence, soon pale green and lustrous, light red or orange-red in their first winter and light or dark brown the following year; usually much smaller. Winter-buds ellipsoidal or ovoid, gradually narrowed at apex, ⅛′—¼′ long, dark reddish brown, and pale-pubescent above the middle. Bark of young stems and branches smooth, light brown, becoming on old trunks ½′—1′ thick and divided by shallow fissures into irregular ridges covered by small light brown scales slightly tinged with red. Wood heavy, hard, strong, coarse-grained, light or reddish brown, with thicker darker colored sapwood.

Distribution. Light dry usually sandy soil; valley of the Androscoggin River, Maine, southern New Hampshire and Vermont to southern Ontario, southward to the District of Columbia and along the Appalachian Mountains to eastern Kentucky and Tennessee, and northern Georgia; in central Georgia and northeastern Mississippi (near Corinth, Alcorn County), and westward through New York, Ohio, Indiana, Illinois and southern Wisconsin to central Missouri (Jerome, Phelps County); in eastern Oklahoma (Arkansas River valley near Fisher, Creek County, G. W. Stevens); ascending to altitudes of nearly 5000° on the southern mountains; the prevailing Oak above 2500° to the summits of the Blue Ridge of the Carolinas; very abundant in the coast region from Massachusetts Bay to southern New Jersey; less common in the interior, growing on dry gravelly uplands, and on the prairies skirting the western margins of the eastern forest.

Occasionally planted in the northeastern states and in Europe as an ornamental tree valued chiefly for the brilliant autumn color of the foliage.

× Quercus Robbinsii Trel., believed to be a hybrid of Quercus coccinea and Q. ilicifolia, occurs at North Easton, Bristol County, Massachusetts.

× Quercus Benderi Baenitz, a supposed hybrid of Quercus coccinea and Q. borealis var. maxima, appeared several years ago in Silesia, and a similar tree has been found in the Blue Hills Reservation near Boston.

6. Quercus palustris Muench. Pin Oak. Swamp Spanish Oak.

Leaves obovate, narrowed and cuneate or broad and truncate at base, divided by wide deep sinuses rounded in the bottom into 5—7 lobes, the terminal lobe ovate, acute, 3-toothed toward the apex, or entire, the lateral lobes spreading or oblique, sometimes falcate, especially those of the lowest pair, gradually tapering and acute at the dentate apex, or obovate and broad at apex, when they unfold light bronze-green stained with red on the margins, lustrous and puberulous above, coated below and on the petioles with pale scurfy pubescence, at maturity thin and firm, dark green and very lustrous above, pale below, with large tufts of pale hairs in the axils of the conspicuous primary veins; 4′—6′ long, 2′—4′ wide, with a stout midrib; late in the autumn gradually turning deep scarlet; petioles slender, yellow, ½′—2′ in length. Flowers: staminate in hairy aments 2′—3′ long; calyx puberulous and divided into 4 or 5 oblong rounded segments more or less laciniately cut on the margins, shorter than the stamens; pistillate on short tomentose peduncles, their involucral scales broadly ovate, tomentose, shorter than the acuminate calyx-lobes; stigmas bright red. Fruit sessile or short-stalked, solitary or clustered; nut nearly hemispheric, about ½′ in diameter, light brown, often striate, inclosed only at the base in a thin saucer-shaped cup dark red-brown and lustrous within, and covered by closely appressed ovate light red-brown thin puberulous scales.

A tree, usually 70°—80° high, with a trunk 2°—3° in diameter, often clothed with small tough drooping branches, or when crowded in the forest sometimes 120° high, with a trunk 60°—70° tall and 4°—5° in diameter, slender branches beset with short-ridged spur-like laterals a few inches in length, forming on young trees a broad pyramidal head, becoming on older trees open and irregular, with rigid and more pendulous branches often furnished at first with small drooping branchlets, and slender tough branchlets dark red and covered by short pale silvery tomentum, soon becoming green and glabrous, lustrous dark red-brown or orange color in their first winter, growing darker in their second year and ultimately dark gray-brown. Winter-buds ovoid, gradually narrowed and acute at apex, about ⅛′ long, with imbricated light chestnut-brown scales puberulous toward the thin sometimes ciliate margins. Bark of young trunks and branches smooth, lustrous, light brown frequently tinged with red, becoming on older trunks ¾′—1¼′ thick, light gray-brown, generally smooth and covered by small closely appressed scales. Wood heavy, hard, strong, coarse-grained, light brown, with thin rather darker colored sapwood; sometimes used in construction, and for shingles and clapboards.

Distribution. Borders of swamps and river-bottoms in deep rich moist soil; valley of the Connecticut River in western Massachusetts and Connecticut; on Grand Isle in the Niagara River, New York to southern Ontario and southwestern Michigan, and westward to eastern Iowa (Muscatine County), and southward to southern West Virginia (Hardy and Mercer Counties), southwestern Virginia (Wythe County), central North Carolina (on Bowling’s Creek, near Chapel Hill, Orange County, and on Dutchman’s Creek, Forsyth County); and to southern Kentucky, central Tennessee, southern Arkansas (Fulton, Hempstead County), and northeastern Oklahoma; rare and of small size in New England; exceedingly common on the coast plain south of the Hudson River; very abundant on the bottom-lands of the streams of the lower Ohio River.

Often cultivated as an ornamental tree in the northeastern states and occasionally in the countries of western and central Europe.

7. Quercus georgiana M. A. Curtis.

Leaves convolute in the bud, elliptic or obovate, gradually narrowed and cuneate at base, divided generally about half way to the midrib by wide or narrow oblique sinuses rounded in the bottom into 3—7 lobes, the terminal lobe ovate, acute, or rounded and entire or frequently furnished with 1 or 2 small lateral teeth, the lateral lobes oblique or spreading, mostly triangular, acute and entire, or those of the upper and of the middle pair often broad and repand-lobulate at the oblique ends, sometimes gradually 3-lobed at the broad apex and narrowed and entire below, or equally 3-lobed, with broad or narrow spreading lateral lobes, or occasionally pinnatifid, when they unfold bright green tinged with red, ciliate on the margins and coated on the midrib, veins, and petioles with loose pale pubescence, at maturity thin, bright green and lustrous above, paler below, and glabrous or furnished with tufts of hairs in the axils of the primary veins, usually about 2½′ long and 1½′ wide; turning dull orange and scarlet in the autumn; petioles slender, ½′—¾′ in length. Flowers: staminate in slender glabrous or pubescent aments 2′—3′ long; calyx divided into 4 or 5 broadly ovate rounded segments rather shorter than the stamens; pistillate on short glabrous slender peduncles; their involucral scales rather shorter than the acute calyx-lobes, pubescent or puberulous; stigmas bright red. Fruit short-stalked; nut ellipsoidal or subglobose, ⅓′—½′ long, light red-brown and lustrous, inclosed for one third to nearly one half its length in a thick cup-shaped cup light red-brown and lustrous on the inner surface, and covered by thin ovate bright light red-brown truncate erose scales.

Distribution. Georgia; on Stone Mountain, and Little Stone Mountain, Dekalb County; on a few other granite hills between the Yellow and Oconee rivers in the region south and east of Stone Mountain (Winder, Jackson County, Rockmart, Polk County and at Warm Springs, Meriwether County).

Occasionally cultivated, and hardy in eastern Massachusetts.

× Quercus Smallii Trel., a possible hybrid of Quercus georgiana and Q. marilandica, occurs on the slopes and summit of Little Stone Mountain, Dekalb County, Georgia.

8. Quercus velutina Lam. Black Oak. Yellow-bark Oak.

Leaves ovate or oblong, rounded, cuneate or truncate at base, mostly 7-lobed and sometimes divided nearly to the middle by wide rounded sinuses into narrow obovate more or less repand-dentate lobes, or into elongated nearly entire mucronate lobes tapering gradually from a broad base, the terminal lobe oblong, elongated, acute, furnished with small lateral teeth, or broad, rounded, and coarsely repand-dentate, or slightly divided into broad dentate lobes or sinuate-dentate, bright crimson when they unfold, and covered above by long loose scattered white hairs and below with thick pale or silvery white tomentum, hoary-pubescent when half grown, and at maturity thick and firm or subcoriaceous, dark green and lustrous above, below yellow-green, brown or dull copper color and more or less pubescent or glabrous with the exception of tufts of rusty hairs in the axils of the principal veins, 3′—12′ long and 2′—10′ wide, but usually 5′—6′ long and 3′—4′ wide, with a stout midrib and primary veins; late in the autumn turning dull red, dark orange color, or brown, and falling gradually during the winter; petioles stout, yellow, glabrous or puberulous, 3′—6′ in length. Flowers: staminate in tomentose or pubescent aments 4′—6′ long; calyx coated with pale hairs, with ovate acute lobes; pistillate on short tomentose peduncles, their involucral scales ovate, shorter than the acute calyx-lobes; stigmas bright red. Fruit sessile or short-stalked, solitary or in pairs; nut ovoid-oblong, obovoid, oval or hemispheric, broad and rounded at base, full and rounded at apex, light red-brown, often striate, frequently coated with soft rufous pubescence, ½′—¾′ long and broad, or rarely 1′ long and broad, inclosed for about half its length or rarely nearly to the apex in the thin deeply cup-shaped or turbinate cup dark red-brown on the inner surface, covered by thin light chestnut-brown acute hoary scales closely appressed at the base of the cup, loosely imbricated above the middle, with free scarious tips forming a fringe-like border to its rim.

A tree, often 70°—80° and occasionally 150° high, with a trunk 3°—4° in diameter, slender branches spreading gradually into a narrow open head, stout branchlets coated at first with pale or fulvous scurfy tomentum, becoming in their first winter glabrous, dull red or reddish brown, growing dark brown in their second year or brown slightly tinged with red. Winter-buds ovoid, strongly angled, gradually narrowed and obtuse at apex, hoary-tomentose, ¼′—½′ long. Bark of young stems and branches smooth, dark brown, deep orange color internally, becoming ¾′—1½′ thick on old trunks, and deeply divided into broad rounded ridges broken on the surface into thick dark brown or nearly black closely appressed plate-like scales. Wood heavy, hard, strong, coarse-grained, bright brown tinged with red, with thin lighter colored sapwood; of little value except as fuel. The bark abounds in tannic acid and is largely used in tanning, as a yellow dye, and in medicine.

Distribution. Dry gravelly uplands and ridges; coast of southern Maine to northern Vermont, southern and western Ontario, the southern peninsula of Michigan, northwestern, eastern and southern Iowa, and southeastern Nebraska, and southward to western Florida, southern Alabama, Mississippi, Louisiana, eastern Kansas, northeastern Oklahoma and eastern Texas to the valley of the Brazos River; one of the commonest Oaks on the gravelly drift of southern New England and the middle states; ascending on the southern Appalachian Mountains to altitudes of about 4000°, and often forming a large part of the forest growth on their foothills; abundant in all parts of the Mississippi basin, and of its largest size in the valley of the lower Ohio River; the common species of the Black Oak group reaching the south-Atlantic and Gulf Coast, and here generally scattered on dry ridges through the maritime Pine belt.

Quercus velutina, which is more variable in the form of its leaves than the other North American Black Oaks, is easily recognized by the bright yellow color of the inner bark, in early spring by the deep red color of the unfolding leaves, becoming pale and silvery in a few days, and by the large tomentose winter-buds. From western Missouri to northwestern Arkansas a form occurs (var. missouriensis Sarg.) with the mature leaves covered above with fascicled hairs, and coated below and on the petioles and summer branchlets with rusty pubescence, and with broader more loosely imbricated hoary-tomentose cup-scales.

9. Quercus Kelloggii Newb. Black Oak.

Quercus californica Coop.

Leaves oblong or obovate, truncate, cuneate or rounded at the narrow base, 7 or rarely 5-lobed by wide and deep or shallow and oblique sinuses rounded in the bottom, the terminal lobe ovate, 3-toothed at the acute apex, the lateral lobes tapering gradually from the base or broad and obovate, coarsely repand-dentate with acute pointed teeth, or rarely entire, when they unfold dark red or purple and pilose above and coated below and on the petioles with thick silvery white tomentum, at maturity thick and firm, lustrous, dark yellow-green and glabrous or rarely pubescent above, light yellow-green or brownish and glabrous or pubescent, or occasionally hoary-tomentose below, 3′—6′ long, 2′—4′ wide; turning yellow or brown in the autumn before falling; petioles slender, yellow, 1′—2′ in length. Flowers: staminate in hairy aments 4′—5′ long; calyx pubescent, divided into 4 or 5 ovate acute segments shorter than the stamens; anthers bright red; pistillate on short tomentose peduncles, their involucral scales ovate, coated like the acute calyx-lobes with pale tomentum; stigmas dark red. Fruit short-stalked, solitary or clustered; nut oblong, ellipsoidal or obovoid, broad and rounded at base, full and rounded or gradually narrowed and acute at the puberulous apex, 1′—1½′ long, about ¾′ broad, light chestnut-brown, often striate, inclosed for one fourth to two thirds of its length in the deep cup-shaped cup light brown on the inner surface, and covered by thin ovate-lanceolate lustrous light chestnut brown scales, sometimes rounded and thickened on the back toward the base of the cup, their tips elongated, thin and erose on the margins, often forming a narrow fringe-like border to the rim of the cup.

A tree, occasionally 100° high, with a trunk 3°—4° in diameter, stout spreading branches forming an open round-topped head, and branchlets coated at first with thick hoary caducous tomentum, bright red or brown tinged with red, and usually glabrous or pubescent or puberulous during their first winter, becoming dark red-brown in their second year; frequently much smaller and at high elevations a small shrub (f. cibata Jeps.). Winter-buds ovoid, gradually narrowed and acute at apex, about ¼′ long, with closely imbricated pale chestnut-brown scales ciliate on the thin scarious margins and pubescent toward the point of the bud. Bark of young stems and branches smooth, light brown, becoming on old trunks 1′—1½′ thick, dark brown slightly tinged with red or nearly black, divided into broad ridges at the base of old trees and broken above into thick irregular oblong plates covered by minute closely appressed scales. Wood heavy, hard, strong, very brittle, bright red, with thin lighter colored sapwood; occasionally used as fuel.

Distribution. Valleys and mountain slopes; basin of the Mackenzie River in western Oregon, southward over the California coast ranges, and along the western slopes of the Sierra Nevada up to altitudes of 6500° to the Cuyamaca Mountains near the southern boundary of California; extending across the Sierra Nevada to the foothills of Owens valley (Jepson) in eastern California; rare in the immediate neighborhood of the coast; the largest and most abundant Oak-tree of the valleys of southwestern Oregon and of the Sierra Nevada, sometimes forming groves of considerable extent in coniferous forests; of its largest size at altitudes of about 6000° above the sea.

10. Quercus Catesbæi Michx. Turkey Oak.

Leaves oblong or obovate or nearly triangular, gradually narrowed and cuneate at base, deeply divided by wide rounded sinuses into 3 or 5 or rarely 7 lobes, the terminal lobe ovate, elongated, acute and entire or repand-dentate, or obovate and coarsely equally or irregularly 3-toothed at apex, the lateral lobes spreading, usually falcate, entire and acute, tapering from the broad base, and broad, oblique, and repand-lobulate at apex, or 3-toothed at the broad apex and gradually narrowed to the base, coated when they unfold with rufous fascicled hairs, and when fully grown thick and rigid, bright yellow-green and lustrous above, paler, lustrous, and glabrous below, with large tufts of rusty hairs in the axils of the veins, 3′—12′ long, 1′—10′ wide, but usually about 5′ long and wide, with a broad yellow or red-brown midrib; turning bright scarlet before falling in the late autumn or early winter; petioles stout, grooved, ¼′—¾′ in length. Flowers: staminate in slender hairy red-stemmed aments 4′—5′ long; calyx puberulous and divided into 4 or 5 ovate acute lobes; pistillate on short stout tomentose peduncles, their involucral scales bright red, pubescent, hairy at the margins; stigmas dark red. Fruit short-stalked, usually solitary; nut oval, full and rounded at the ends, about 1′ long and ¾′ broad, dull light brown, covered at the apex by a thin coat of snow-white tomentum, inclosed for about one third its length in a thin turbinate cup often gradually narrowed into a stout stalk-like base, light red-brown and lustrous on the inner surface, covered by ovate-oblong rounded scales extending above the rim of the cup and down over the upper third of the inner surface, and hoary-pubescent except their thin bright red margins.

A tree, usually 20°—30°, or occasionally 50°—60° high, with a trunk rarely exceeding 2° in diameter, stout spreading more or less contorted branches forming a broad or narrow open irregular generally round-topped head, and stout branchlets coated at first with fascicled hairs, nearly glabrous and deep red when the leaves are half grown, dark red in their first winter, gradually growing dark brown; generally much smaller and sometimes shrubby. Winter-buds elongated, acute, ½′ long, with light chestnut-brown scales erose on the thin margins, and coated, especially toward the point of the bud, with rusty pubescence. Bark ½′—1′ thick, red internally, dark gray tinged with red on the surface, and at the base of old trunks becoming nearly black, deeply and irregularly furrowed and broken into small appressed scales. Wood heavy, hard, strong, rather close-grained, light brown tinged with red, with thick lighter colored sapwood; largely used for fuel.

Distribution. Dry barren sandy ridges and sandy bluffs and hummocks in the neighborhood of the coast; southeastern Virginia (near Zuni, Isle of Wight County) to the shores of Indian River and Peace Creek, Florida, and westward to eastern Louisiana; comparatively rare toward the western limits of its range, and most abundant and of its largest size on the high bluff-like shores of bays and estuaries in South Carolina and Georgia; the prevailing tree with Quercus cinerea in the flat woods of the interior of the Florida peninsula as far south as the sandy ridges in the neighborhood of Lake Istokpoga, De Soto County.

× Quercus Mellichampii Trel. believed to be a hybrid of Quercus Catesbæi and Q. laurifolia occurs at Bluffton on the coast of South Carolina, in the neighborhood of Orlando, Orange County and near San Mateo, Putnam County, Florida.

× Quercus Ashei Trel. believed to be a hybrid of Quercus Catesbæi with Q. cinerea occurs at Folkston and near Trader’s Hill, Charlton County and St. Mary’s, Camden County, Georgia.

× Quercus blufftonensis Trel., a probable hybrid of Quercus Catesbæi and Q. rubra L., has been found at Bluffton, South Carolina.

× Quercus Walteriana Ashe, believed to be a hybrid of Quercus Catesbæi and Q. nigra, is not rare in the immediate neighborhood of the coast of South Carolina and Georgia, and occurs on sand hills in Sampson County, North Carolina, near Jacksonville, Duval County, Florida, at Mount Vernon, Mobile County and in the neighborhood of Selma, Dallas County, Alabama.

11. Quercus ilicifolia Wang. Bear Oak. Scrub Oak.

Quercus nana Sarg.

Leaves obovate or rarely oblong, gradually or abruptly cuneate at base, divided by wide shallow sinuses into 3—7, usually 5, acute lobes, the terminal lobe ovate, elongated, rounded and 3-toothed or acute and dentate or entire at apex, the lateral lobes spreading, mostly triangular and acute, or those of the upper pair broad, oblique and repand-lobulate or broad at apex, slightly 3-lobed and entire below, or deeply 3-lobed above and sinuate below, or occasionally oblong to oblong-obovate and entire, with undulate margins, when they unfold dull red and puberulous or pubescent on the upper surface and coated on the lower and on the petioles with thick pale tomentum, with conspicuous tufts of silvery white hairs in the axils of the veins, at maturity thick and firm, dark green and lustrous above, covered below with pale or silvery white pubescence, 2′—5′ long, 1½′—3′ wide, with a stout yellow midrib and slender primary veins; turning dull scarlet or yellow in the autumn; petioles slender, glabrous, or pubescent, 1′—1½′ in length. Flowers: staminate in hairy aments 4′—5′ long, and often persistent until midsummer; calyx red or green tinged with red and irregularly divided into 3—5 ovate rounded lobes shorter than the stamens; anthers bright red ultimately yellow; pistillate on stout tomentose peduncles, their involucral scales ovate, about as long as the acute calyx-lobes, red and tomentose; stigmas dark red. Fruit produced in great profusion, sessile or stalked, in pairs or rarely solitary; nut ovoid, broad, flat or rounded at base, gradually narrowed and acute or rounded at apex, about ½′ long and broad, light brown, lustrous, usually faintly striate, inclosed for about one half its length in the cup-shaped or saucer-shaped cup often abruptly enlarged above the stalk-like base, thick, light reddish brown within, and covered by thin ovate closely imbricated red-brown puberulous scales acute or truncate at apex, the minute free tips of the upper scales forming a fringe-like border to the cup.

A tree, occasionally 18°—20° high, with a trunk 5′—6′ in diameter, with slender spreading branches usually forming a round-topped head, and slender branchlets dark green more or less tinged with red and hoary-pubescent at first, during their first winter red-brown or ashy gray and pubescent or puberulous, becoming glabrous and darker in their second year and ultimately dark brown or nearly black; more frequently an intricately branched shrub, with numerous contorted stems 3°—10° tall. Winter-buds ovoid, obtuse, about ⅛′ long, with dark chestnut-brown rather loosely imbricated glabrous or pilose scales. Bark thin, smooth, dark brown, covered by small closely appressed scales.

Distribution. Dry sandy barrens and rocky hillsides; coast of eastern Maine southward through eastern and southern New England to southern and southwestern Pennsylvania and along the Appalachian Mountains, principally on their eastern slopes, to southern Virginia; on Crowder and King Mountains, Gaston County, North Carolina; and westward to the shores of Lake George and the valley of the Hudson River; common in eastern and southern New England, in the Pine barrens of New Jersey, and in eastern Pennsylvania.

× Quercus Brittonii Davis, believed to be a hybrid of Quercus ilicifolia and Q. marilandica, has been found on Staten Island, New York, and at Ocean Grove, Monmouth County, New Jersey.

× Quercus Giffordii Trel., believed to be a hybrid of Quercus ilicifolia and Q. Phellos, has been found at May’s Landing, Atlantic County, New Jersey.

× Quercus Rehderi Trel., believed to be a hybrid of Quercus ilicifolia and Q. velutina, is not rare in eastern Massachusetts and occurs on Martha’s Vineyard (Chilmark).

12. Quercus rubra L. Red Oak. Spanish Oak.

Quercus digitata Sudw.

Leaves ovate to obovate, narrowed and rounded or cuneate at base, the terminal lobe long-acuminate, entire or slightly lobed, often falcate, usually longer than the 2 or 4 acuminate entire lateral lobes narrowed from a broad base and often falcate, or oblong-obovate and divided at the broad apex by wide or narrow sinuses broad and rounded in the bottom into 3 rounded or acute entire or dentate lobes, and entire and gradually narrowed below into an acute or rounded base (var. triloba Ashe), the two forms usually occurring on different but sometimes on the same tree, at maturity thin and firm, dark green and lustrous above, coated below with soft close pale or rusty pubescence, 6′—7′ long and 4′—5′ wide, obscurely reticulate-venulose, with a stout tomentose midrib and primary veins; turning brown or dull orange color in the autumn; petioles slender, flattened, 1′—2′ in length. Flowers: staminate in tomentose aments, 3′—5′ long; calyx thin and scarious, pubescent on the outer surface, divided into 4 or 5 ovate rounded segments; pistillate on stout tomentose peduncles, their involucral scales coated with rusty tomentum, as long or rather shorter than the acute calyx-lobes; stigmas dark red. Fruit sessile or short-stalked; nut subglobose to ellipsoidal, full and rounded at apex, truncate and rounded at base, about ½′ long, bright orange-brown, inclosed only at base or sometimes for one third its length in a thin saucer-shaped cup flat on the bottom or gradually narrowed from a stalk-like base, or deep and turbinate, bright red-brown on the inner surface, covered by thin ovate-oblong reddish scales acute or rounded at apex and pale-pubescent except on the margins.

A tree, usually 70°—80° high, with a trunk 2°—3° in diameter, large spreading branches forming a broad round-topped open head, and stout branchlets coated at first, like the young leaves, with thick rusty or orange-colored clammy tomentum, dark red or reddish brown and pubescent or rarely glabrous during their first winter, becoming in their second year dark red-brown or ashy gray. The var. triloba usually 20°—30° rarely 40°—50° high. Winter-buds ovoid or oval, acute, ⅛′—¼′ long, with bright chestnut-brown puberulous or pilose scales ciliate with short pale hairs. Bark ¾′—1′ thick, dark brown or pale, and divided by shallow fissures into broad ridges covered by thin closely appressed scales. Wood hard, strong, not durable, coarse-grained, light red, with thick lighter colored sapwood; sometimes used in construction, and largely as fuel. The bark is rich in tannin, and is used in tanning leather and occasionally in medicine.

Distribution. Southeastern and southern Pennsylvania and southern New Jersey southward to central Florida, through the Gulf states to the valley of the Brazos River, Texas, and through eastern Oklahoma and southwestern Missouri to central Tennessee and Kentucky, southern Indiana and Illinois, southern Ohio (Black Fork Creek, Lawrence County), and Kanawha County, West Virginia; in the north Atlantic states only in the neighborhood of the coast and comparatively rare; very common in the south Atlantic and Gulf states on dry hills between the coast plain and the Appalachian Mountains; less abundant in the southern maritime Pine belt. The var. triloba: rare and local. Pleasant Grove, Lancaster County, Pennsylvania and Jefferson County, Indiana, southward to central and western Florida, southern Alabama and Mississippi, western Arkansas and eastern Texas; on dry uplands near Milledgeville, Baldwin County, Georgia, the prevailing form.

Quercus rubra var. pagodæfolia Ashe. Swamp Spanish Oak. Red Oak.

Quercus pagoda Rafn.
Quercus pagodæfolia Ashe.

Leaves elliptic to oblong, acuminate, gradually narrowed and cuneate or full and rounded or rarely truncate at base, deeply divided by wide sinuses rounded in the bottom into 5—11 acuminate usually entire repand-dentate lobes often falcate and spreading at right angles to the midrib or pointed toward the apex of the leaf, when they unfold coated with pale tomentum, thickest on the lower surface, and dark red on the upper surface, at maturity dark green and very lustrous above, pale and tomentose below, 6′—8′ long and 5′—6′ wide, with a stout midrib usually puberulous on the upper side, slender primary veins arched to the points of the lobes, and conspicuous reticulate veinlets; turning bright clear yellow before falling; petioles stout, pubescent or tomentose, 1½′—2′ in length. Flowers and Fruit as in the species.

A tree, sometimes 120° high, with a trunk 4°—5° in diameter, heavy branches forming in the forest a short narrow crown, or in more open situations wide-spreading or ascending and forming a great open head, and slender branchlets hoary tomentose at first, tomentose or pubescent during their first winter, and dark reddish brown and puberulous during their second year. Winter-buds ovoid, acute, often prominently 4-angled, about ¼′ long, with light red-brown puberulous scales sometimes ciliate at the apex. Bark about 1′ thick and roughened by small rather closely appressed plate-like light gray, gray-brown or dark brown scales. Wood light reddish brown, with thin nearly white sapwood; largely manufactured into lumber in the Mississippi valley, and valued almost as highly as white oak.

Distribution. Rich bottom-lands and the alluvial banks of streams; Maryland (Queen Anne County) and coast of Virginia to northern Florida, and through the Gulf states and Arkansas to southern Missouri, western Tennessee and Kentucky, and southern Illinois and Indiana; most abundant and one of the largest and most valuable timber-trees in the river swamps of the Yazoo basin, Mississippi, and of eastern Arkansas. Differing chiefly from the type in the more numerous and more acuminate lobes of the usually more elongated leaves usually paler on the lower surface, and in the generally paler bark of the trunk; passing into Quercus rubra var. leucophylla Ashe with leaves on upper branches nearly as broad as long thickly covered below with brownish pubescence and deeply divided into 5—7 lobes, and on lower branches slightly obovate, less deeply divided, thin, dark green, sometimes pubescent becoming glabrous above and often covered below with pale or brown pubescence.

A tree sometimes 120° high; in low rich soil; coast region of southeastern Virginia, southward to western Florida and through the Gulf states to the valley of the Neches River, Texas, and northward to northern Arkansas; in southern Illinois (near Mt. Carmel, Wabash County) and southwestern Indiana (near Hovey Lake, Posey County); abundant in low woods about River Junction, Gadsden County, Florida, and in central Mississippi.

× Quercus Willdenoviana Zabel is believed in Europe to be a hybrid of Quercus rubra and Quercus velutina.

13. Quercus marilandica Muench. Black Jack. Jack Oak.

Leaves broadly obovate, rounded or cordate at the narrow base, usually 3 or rarely 5-lobed at the broad and often abruptly dilated apex, with short or long, broad or narrow, rounded or acute, entire or dentate lobes, or entire or dentate at apex, sometimes oblong-obovate, undulate-lobed at the broad apex and entire below, or equally 3-lobed with elongated spreading lateral lobes broad and lobulate at apex, when they unfold coated with a clammy tomentum of fascicled hairs and bright pink on the upper surface, at maturity thick and firm or subcoriaceous, dark yellow-green and very lustrous above, yellow, orange color, or brown and scurfy-pubescent below, usually 6′—7′ long and broad, with a thick broad orange-colored midrib; turning brown or yellow in the autumn; petioles stout, yellow, glabrous or pubescent, ½′—¾′ in length. Flowers: staminate in hoary aments 2′—4′ long; calyx thin and scarious, tinged with red above the middle, pale-pubescent on the outer surface, divided into 4 or 5 broad ovate rounded lobes; anthers apiculate, dark red; pistillate on short rusty-tomentose peduncles coated like their involucral scales with thick rusty tomentum; stigmas dark red. Fruit, solitary or in pairs, usually pedunculate; nut oblong, full and rounded at the ends, rather broader below than above the middle, about ¾′ long, light yellow-brown and often striate, the shell lined with dense fulvous tomentum, inclosed for one third to nearly two thirds of its length in a thick turbinate light brown cup puberulous on the inner surface, and covered by large reddish brown loosely imbricated scales often ciliate and coated with loose pale or rusty tomentum, the upper scales smaller, erect, inserted on the top of the cup in several rows, and forming a thick rim round its inner surface, or occasionally reflexed and covering the upper half of the inner surface of the cup.

A tree, 20°—30°, or occasionally 40°—50° high, with a trunk rarely more than 1′ in diameter, short stout spreading often contorted branches forming a narrow compact round-topped or sometimes an open irregular head, and stout branchlets coated at first with thick pale tomentum, light brown and scurfy-pubescent during their first summer, becoming reddish brown and glabrous or puberulous in the winter, and ultimately brown or ashy gray. Winter-buds ovoid or oval, prominently angled, light red-brown, coated with rusty brown hairs, about ¼′ long. Bark 1′—1½′ thick, and deeply divided into nearly square plates 1′—3′ long and covered by small closely appressed dark brown or nearly black scales. Wood heavy, hard, strong, dark rich brown, with thick lighter colored sapwood; largely used as fuel and in the manufacture of charcoal.

Distribution. Dry sandy or clay barrens; Long Island and Staten Island, New York, eastern and southern Pennsylvania, and southern New Jersey to the shores of Matanzas Inlet and Tampa Bay, Florida, and westward through the Gulf states to western Texas (Callahan County) and to western Oklahoma (Dewey and Kiowa Counties), Arkansas, eastern Kansas, southeastern Nebraska and through Missouri to northeastern Illinois, southwestern and southern Indiana, and northeastern Kentucky (South Portsmouth, Greenup County, R. E. Horsey); rare in the north, very abundant southward; west of the Mississippi River often forming on sterile soils a great part of the forest growth; of its largest size in southern Arkansas and eastern Texas.

× Quercus Rudkinii Britt., with characters intermediate between those of Quercus marilandica and Q. Phellos, and probably a hybrid of these species, has been found near Tottenville, Staten Island, New York, at Keyport, Monmouth County, New Jersey, and at the Falls of the Yadkin River, Stanley County, North Carolina.

× Quercus sterilis Trel., believed to be a hybrid of Quercus marilandica and Q. nigra has been found in Bladen County, North Carolina.

× Quercus Hastingsii Sarg., believed to be a hybrid of Quercus marilandica and Q. texana, occurs near Boerne, Kendall County, and at Brownwood, Brown County, Texas.

× Quercus Bushii Sarg., believed to be a hybrid of Quercus marilandica and Q. velutina, although not common, occurs in eastern Oklahoma (Sapulpa, Creek County), Mississippi (Oxford, Lafayette County), Alabama (Dothan, Houston County, near Berlin, Dallas County, and Daphne, Baldwin County), Florida (Sumner, Levey County), and in Georgia (Climax, Decatur County).

14. Quercus arkansana Sarg.

Leaves broadly obovate, slightly 3-lobed or dentate at the wide apex, cuneate at base, on sterile branches often oblong-ovate, acute or rounded at apex, rounded at base, the lobes ending in long slender mucros, when they unfold tinged with red, thickly covered with pale fascicled hairs persistent until summer, the midrib and veins more thickly clothed with long straight hairs, and at maturity glabrous, with the exception of small axillary tufts of pubescence on the lower surface, light yellow-green above, paler below, 2′—2¾′ long and broad, with a slender light yellow midrib, thin primary veins and prominent veinlets; on sterile branches often 4½′—5½′ long and 2½′—2¾′ wide; petioles slender, coated at first with clusters of pale hairs, becoming glabrous or puberulous, ⅗′—⅘′ in length. Flowers: staminate in aments covered with clusters of long pale hairs, 2′—2½′ long; calyx usually 4 rarely 3-lobed, thinly covered with long white hairs; stamens usually 4; anthers ovoid-oblong, apiculate, dark red; pistillate on stout peduncles, hoary-tomentose like the scales of the involucre; stigmas dark red. Fruit solitary or in pairs, on short glabrous peduncles; nut broad-ovoid, rounded at apex, sparingly pubescent especially below the middle with fascicled hairs, light brown, obscurely striate, ¼′—⅓′ long, ½′—⅝′ thick, inclosed only at base in the flat saucer-shaped cup, pubescent on the inner surface, covered with closely appressed scales obtuse at their narrow apex, red on the margins, pale pubescent, those of the upper rank smaller, erect, inserted on the top of the cup and forming a thin rim round its inner surface.

A tree when crowded in the forest often 60°—70° high, with a tall trunk, stout ascending branches forming a long narrow head, and slender branchlets thickly coated early in the season with pale fascicled hairs, pubescent or nearly glabrous in their first autumn and darker and glabrous in their second year, when not crowded by other trees rarely 40° high with a short trunk occasionally 1° in diameter. Winter-buds ovoid, acute, with thin light chestnut-brown slightly pubescent or nearly glabrous scales. Bark thick, nearly black, divided by deep fissures into long narrow ridges covered with thick closely appressed scales.

Distribution. Low woods and on rolling sand hills four miles north of Fulton, Hempstead County, Arkansas; rare and local.

15. Quercus nigra L. Water Oak.

Leaves oblong-obovate, gradually narrowed and cuneate at base and enlarged often abruptly at the broad rounded entire or occasionally 3-lobed apex, on vigorous young branchlets sometimes pinnatifid with acute, acuminate or rounded lobes or broadly oblong-obovate and rounded at apex with entire or undulate margins, on upper branches occasionally linear-lanceolate, on occasional trees narrowed below to an elongated cuneate base and gradually widened above into a more or less deeply 3-lobed apex, the lobes rounded or acute (var. tridentifera Sarg.), or often acute at the ends, and on upper branchlets sometimes linear-lanceolate to linear-obovate, acute or rounded at apex, divided above the middle by deep wide rounded sinuses into elongated lanceolate acute entire lobes, or pinnatifid above the middle, when they unfold thin, light green more or less tinged with red and covered by fine caducous pubescence, with conspicuous tufts of pale hairs in the axils of the veins below, at maturity thin, dull bluish green, paler below than above, glabrous or with axillary tufts of rusty hairs, usually about 2½′ long and 1½′ wide, or on fertile branches sometimes 6′ long and 2½′ wide; turning yellow and falling gradually during the winter; petioles stout, flattened, ⅛′—½′ in length; leaves of seedling plants linear-lanceolate with entire or undulate margins, or occasionally lobed with 1 or 2 pointed lobes, often deeply 3-lobed at a wide apex, and occasionally furnished below the middle with a single acuminate lobe, all the forms often occurring on a plant less than three feet high. Flowers: staminate in red hairy-stemmed aments 2′—3′ long; calyx thin and scarious, covered on the outer surface with short hairs, divided into 4 or 5 ovate rounded segments; pistillate on short tomentose peduncles, their involucral scales a little shorter than the acute calyx-lobes and coated with rusty hairs; stigmas deep red. Fruit usually solitary, sessile or short-stalked; nut ovoid, broad and flat at base, full and rounded at the pubescent apex, light yellow-brown, often striate, ⅓′—⅔′ long and nearly as thick, usually inclosed only at the base in a thin saucer-shaped cup, or occasionally for one third its length in a cup-shaped cup, coated on the inner surface with pale silky tomentum and covered by ovate acute closely appressed light red-brown scales clothed with pale pubescence except on their darker colored margins.

A tree, occasionally 80° high, with a trunk 2°—3½° in diameter, numerous slender branches spreading gradually from the stem and forming a symmetrical round-topped head, and slender glabrous branchlets light or dull red during their first winter, becoming grayish brown in their second season. Winter-buds ovoid, acute, strongly angled, covered by loosely imbricated dark red-brown puberulous scales slightly ciliate on the thin margins. Bark ½′—¾′ thick, with a smooth light brown surface slightly tinged with red and covered by smooth closely appressed scales. Wood heavy, hard, strong, close-grained, light brown, with thick lighter colored sapwood; little valued except as fuel.

Distribution. High sandy borders of swamps and streams and the rich bottom-lands of rivers, or northward sometimes in dry woods; southern Delaware, southward to the shores of the Indian River and Tampa Bay, Florida, ranging inland in the south Atlantic states through the Piedmont region, and westward through the Gulf states to the valley of the Colorado River, Texas, and through eastern Oklahoma and Arkansas to southeastern Missouri and to central Tennessee and Kentucky. The var. tridentifera Sarg. rare and local; southwest Virginia to Alabama (near Selma, Dallas County), central and western Mississippi, eastern Louisiana; valley of Navidad River, Lavaca County, Texas. A form (f. microcarya Sarg.—Quercus microcarya Small) occurs in the dry soil on slopes of Little Stone Mountain, Dekalb County, Georgia.

The Water Oak is commonly planted as a shade-tree in the streets and squares of the cities and towns of the southern states.

16. Quercus rhombica Sarg.

Leaves rhombic, rarely oblong-obovate to lanceolate, acute or rounded and apiculate at apex, cuneate at base, the margins entire or slightly undulate, those on vigorous shoots occasionally furnished on each side near the middle with a short lobe, when they unfold deeply tinged with red, covered with short pale caducous pubescence and furnished below with usually persistent tufts of axillary hairs, at maturity thin, dark green and lustrous above, pale below, 3½′—4′ long, 1½′—2′ wide, with a stout conspicuous yellow midrib and slender forked primary veins; turning yellow and falling gradually in early winter, rarely at the ends of branches, obovate and rounded, slightly 3-lobed or undulate at the broad apex (var. obovatifolia Sarg.); petioles yellow, ⅕′—½′ in length. Flowers not seen. Fruit sessile or short-stalked; nut ovoid, rounded at apex, thickly covered with pale pubescence, ⅖′—½′ long, ⅗′ thick; inclosed only at the base in a saucer-shaped cup, rounded on the bottom, silky pubescent on the inner surface, and covered with slightly pubescent reddish brown loosely appressed scales rounded at apex, with free tips, those of the upper rank thin and ciliate on the margins.

A tree often 120°—150° high, with a tall trunk 3°—4½° in diameter, stout, wide-spreading smooth branches forming a broad open head, and slender glabrous branchlets red-brown during their first season and dark gray the following year. Bark pale gray, slightly furrowed and covered with closely appressed scales, ½′—¾′ thick.

Distribution. Borders of swamps and low wet woods of the coast region; southeastern Virginia (Dismal Swamp) to northern Florida, and through the Gulf states to the valley of the Neches River (Beaumont, Jefferson County), eastern Texas; in Louisiana northward to the valley of the Red River; most abundant in south central Alabama and in Louisiana.

× Quercus beaumontiana Sarg., believed to be a hybrid of Quercus rhombica and Q. rubra has been found growing by a street in Beaumont, Jefferson County, Texas.

× Quercus Cocksii Sarg., probably a hybrid of Quercus rhombica and Q. velutina, has been found at Pineville, Rapides Parish, Louisiana.

17. Quercus Phellos L. Willow Oak.

Leaves ovate-lanceolate or rarely obovate-lanceolate, often somewhat falcate, gradually narrowed and acute at the ends, and entire with slightly undulate margins, when they fold light yellow-green and lustrous on the upper surface, coated on the lower with pale caducous pubescence, at maturity glabrous, light green and rather lustrous above, dull and paler or rarely hoary-pubescent below, conspicuously reticulate-venulose, 2½′—5′ long, ¼′—1′ wide, with a slender yellow midrib and obscure primary veins forked and united about halfway between the midrib and margins; turning pale yellow in the autumn; petioles stout, about ⅛′ in length. Flowers: staminate in slender-stemmed aments 2′—3′ long; calyx yellow, hirsute, with 4 or 5 acute segments; pistillate on slender glabrous peduncles, their involucral scales brown covered by pale hairs, about as long as the acute calyx-lobes; stigmas bright red. Fruit short-stalked or nearly sessile, solitary or in pairs; nut hemispheric, light, yellow-brown, coated with pale pubescence, inclosed only at the very base in the thin pale reddish brown saucer-shaped cup silky-pubescent on the inner surface, and covered by thin ovate hoary-pubescent closely appressed scales rounded at apex.

A tree, often 70°—90° high, with a trunk 2° or rarely 4° in diameter, small branches spreading into a comparatively narrow open or conical round-topped head, and slender glabrous reddish brown branchlets roughened by dark lenticels, becoming in their second year dark brown tinged with red or grayish brown; usually much smaller. Winter-buds ovoid, acute, about ⅛′ long, with dark chestnut-brown scales pale and scarious on the margins. Bark ½′—¾′ thick, light red-brown slightly tinged with red, generally smooth but on old trees broken by shallow narrow fissures into irregular plates covered by small closely appressed scales. Wood heavy, strong, not hard, rather coarse-grained, light brown tinged with red, with thin lighter colored sapwood; occasionally used in construction, for clapboards and the fellies of wheels.

Distribution. Low wet borders of swamps and streams and rich sandy uplands; Staten Island, New York, southern New Jersey and southeastern Pennsylvania and southward to northeastern Florida, through the Gulf states to the valley of the Navasota River, Brazos County, Texas, and through Arkansas, eastern Oklahoma and southeastern Missouri to central Tennessee and northwestern Kentucky (Ballard County), and in southwestern Illinois (Massac and Pope Counties); in the Atlantic states usually confined to the maritime plain; less common in the middle districts, rarely extending to the Appalachian foothills.

Occasionally planted as a shade-tree in the streets of southern towns, and rarely in western Europe; hardy in eastern Massachusetts.

Quercus heterophylla Michx. f.

This has usually been considered a hybrid between Quercus Phellos and Quercus velutina or Quercus borealis var. maxima; first known in the eighteenth century from an individual growing in a field belonging to John Bartram on the Schuylkill River, Philadelphia. What appears to be the same form has since been discovered in a number of stations from New Jersey to Texas, and it is possible that Quercus heterophylla may, as many botanists have believed, best be considered a species.

× Quercus subfalcata Trel., believed to be a hybrid of Quercus Phellos and Q. rubra has been found at Wickliffe, Ballard County, Illinois, at Campbell, Lawrence County, Mississippi, Fulton, Hempstead County, Arkansas, and Houston, Harris County, Texas; its var. microcarpa Sarg., probably of the same parentage, originated in a Dutch nursery.

× Quercus ludoviciana Sarg., believed to be a hybrid of Quercus Phellos and Q. rubra var. pagodæfolia grows in low wet woods ten miles west of Opelousas, St. Landry Parish, Louisiana.

18. Quercus laurifolia Michx. Laurel Oak. Water Oak.

Leaves elliptic or rarely slightly broadest above the middle, acuminate at the ends, apiculate at apex, occasionally lanceolate or oblong-obovate and rounded at apex (var. hybrida Michx.) sometimes 3-lobed at apex, the terminal lobe acuminate, much larger than the others (var. tridentata Sarg.), frequently unequally lobed on vigorous branches of young trees, with small nearly triangular lobes, when they unfold in spring yellow-green, or later in the season often pink or bright red, and slightly puberulous, at maturity thin, green, and very lustrous above, light green and less lustrous below, usually 3′—4′ long and ¾′ wide, with a conspicuous yellow midrib; falling abruptly in early spring leaving the branches bare during only a few weeks; petioles stout, yellow, rarely more than ¼′ in length. Flowers: staminate in red-stemmed hairy aments 2′—3′ long; calyx pubescent on the outer surface, divided into 4 ovate rounded lobes; pistillate on stout glabrous peduncles, their involucral scales brown and hairy, about as long as the acute calyx-lobes; stigmas dark red. Fruit sessile or subsessile, generally solitary; nut ovoid to hemispheric, broad and slightly rounded at base, full and rounded at the puberulous apex, dark brown, about ½′ long, inclosed for about one fourth its length in a thin saucer-shaped cup red-brown and silky-pubescent on the inner surface, and covered by thin ovate light red-brown scales rounded at apex and pale-pubescent except on their darker colored margins.

A tree, occasionally 100° high, with a tall trunk 3°—4° in diameter, and comparatively slender branches spreading gradually into a broad dense round-topped shapely head, and slender glabrous branchlets dark red when they first appear, dark red-brown during their first winter, becoming reddish brown or dark gray in their second season. Winter-buds broadly ovoid or oval, abruptly narrowed and acute at apex, /16′—⅛′ long with numerous thin closely imbricated bright red-brown scales ciliate on the margins. Bark of young trees ½′—1′ thick, dark brown more or less tinged with red, roughened by small closely appressed scales, becoming at the base of old trees 1′—2′ thick, nearly black, and divided by deep fissures into broad flat ridges. Wood heavy, very strong and hard, coarse-grained, liable to check badly in drying, dark brown tinged with red, with thick lighter colored sapwood; probably used only as fuel.

Distribution. Sandy banks of streams and swamps and rich hummocks in the neighborhood of the coast; North Carolina (near Newbern) southward to the shores of Bay Biscayne and the valley of the Caloosahatchie River, Florida, and in the interior of the peninsula to the neighborhood of Lake Istokpaga, De Soto County, and westward to eastern Louisiana, ranging inland to Darlington, Darlington County, South Carolina, to the neighborhood of Augusta, Richmond County, Mayfield, Hancock County, Albany, Dougherty County, Cuthbert, Randolph County, and Bainbridge, Decatur County, Georgia, Georgiana, Butler County, and Berlin, Dallas County, Alabama, Rockport, Copiah County, Mississippi, and to the neighborhood of Bogalusa, Washington Parish, Louisiana (R. S. Cocks); nowhere abundant, but most common and of its largest size in eastern Florida.

19. Quercus cinerea Michx. Blue Jack. Upland Willow Oak.

Quercus brevifolia Sarg.

Leaves oblong-lanceolate to oblong-obovate, gradually narrowed and cuneate or sometimes rounded at base, acute or rounded and apiculate at apex, entire with slightly thickened undulate margins, or at the ends of vigorous sterile branches occasionally 3-lobed at the apex and variously lobed on the margins (β dentato-lobata A. De Candolle), when they unfold bright pink and pubescent on the upper surface, coated on the lower with thick silvery white tomentum, at maturity firm in texture, blue-green, lustrous, conspicuously reticulate venulose above, pale-tomentose below, 2′—5′ long, ½′—1½′ wide, with a stout yellow midrib and remote obscure primary veins forked and united within the margins; turning red and falling gradually late in the autumn or in early winter; petioles stout, ¼′—½′ in length. Flowers: staminate in hoary-tomentose aments 2′—3′ long; calyx pubescent, bright red and furnished at apex with a thick tuft of silvery white hairs before opening, divided into 4 or 5 ovate acute lobes, becoming yellow as it opens; stamens 4 or 5; anthers apiculate, dark red in the bud, becoming yellow; pistillate on short stout tomentose peduncles, their involucral scales about as long as the acute calyx-lobes and coated with pale tomentum; stigmas dark red. Fruit produced in great profusion, sessile or raised on a short stalk rarely ¼′ long; nut ovoid, full and rounded at the ends or subglobose, about ½′ long, often striate, hoary-pubescent at apex, inclosed only at the base or for one half its length in a thin saucer-shaped or cup-shaped cup bright red-brown and coated with lustrous pale pubescence on the inner surface, and covered by thin closely imbricated ovate-oblong scales hoary-tomentose except on the dark red-brown margins.

A tree on dry hills, usually 15°—20° high, with a trunk 5′—6′ in diameter, stout branches forming a narrow irregular-head, and thick rigid branchlets coated at first with a dense fulvous or hoary tomentum of fascicled hairs, soon becoming glabrous or puberulous, dark brown sometimes tinged with red during their first winter and darker in their second year; or in low moist soil often 60°—75° high, with a trunk 18′—20′ in diameter, and a broad round-topped shapely head of drooping branches. Winter-buds ovoid, acute, with numerous rather loosely imbricated bright chestnut-brown scales ciliate on the margins, often ¼′ long on vigorous branches, frequently obtuse and occasionally much smaller. Bark ¾′—1½′ thick, and divided into thick nearly square plates 1′—2′ long, and covered by small dark brown or nearly black scales slightly tinged with red. Wood hard, strong, close-grained, light brown tinged with red, with thick darker colored sapwood; probably only used as fuel.

Distribution. Sandy barrens and dry upland ridges, and in the rich moist soil of the pine-covered flats of the Florida peninsula; North Carolina southward to the shores of the Indian River and Peace Creek, Florida, and along the Gulf coast to the valley of the Brazos River, Texas; in the Atlantic and middle Gulf states mostly confined to a maritime belt 40°—60° wide, extending across the Florida peninsula as far south as the sand hills in the neighborhood of Lake Istokpoga, De Soto County, and west of the Mississippi River, ranging inland to the neighborhood of Dallas, Dallas County, Texas and to southeastern Oklahoma (near Antlers, Pushmataha County).

× Quercus dubia Ashe, believed to be a hybrid of Quercus cinerea and Q. laurifolia occurs at Abbottsburg, Bladen County, North Carolina, on the coast of South Carolina, in southern Georgia and northern and central Florida, and at Mississippi City, Lincoln County, Mississippi.

× Quercus subintegra Trel., a supposed hybrid of Quercus cinerea and Q. rubra occurs at Lumber City, Telfair County, Georgia, Lake City, Columbia County, Florida, and at Berlin, Dallas County, Alabama.

× Quercus sublaurifolia Trel., a supposed hybrid of Quercus cinerea and Q. laurifolia occurs at Folkston, Charlton County, Georgia, and at Biloxi, Harrison County, Mississippi.

× Quercus carolinensis Trel., believed to be a hybrid of Quercus cinerea and Q. marilandica occurs at Newbern, Craven County, North Carolina, Lumber City, Telfair County and Climax, Decatur County, Georgia, and near Fletcher, Hardin County, Texas.

× Quercus caduca Trel., believed to be a hybrid of Quercus cinerea and Q. nigra, occurs at Folkston, Charlton County and Lumber City, Telfair County, Georgia, Jacksonville, Duval County, and Gainsville, Alachua County, Florida, Mississippi City, Harrison County, Mississippi, and at Milano, Milano County and Bryan, Brazos County, Texas.

× Quercus oviedoensis Sarg., believed to be a hybrid of Quercus cinerea and Q. myrtifolia, has been found near Oviedo, Orange County, Florida.

20. Quercus imbricaria Michx. Shingle Oak. Laurel Oak.

Leaves oblong-lanceolate to oblong-obovate, apiculate and acute or rounded at apex, gradually narrowed and cuneate or rounded at base, entire with slightly thickened, revolute often undulate margins, or sometimes more or less 3-lobed, or on sterile branches occasionally repand-lobulate, when they unfold bright red, soon becoming yellow-green, covered with scurfy rusty pubescence on the upper surface and hoary-tomentose on the lower, at maturity thin, glabrous, dark green, and very lustrous above, pale green or light brown and pubescent below, 4′—6′ long, ¾′—2′ wide, with a stout yellow midrib, numerous slender yellow veins arcuate and united at some distance from the margins, and reticulate veinlets; late in the autumn turning dark red on the upper surface; petioles stout, pubescent, rarely more than ½′ in length. Flowers: staminate in hoary-tomentose aments, 2′—3′ long; calyx light yellow, pubescent, and divided into 4 acute segments; pistillate on slender tomentose peduncles, their involucral scales covered with pale pubescence and about as long as the acute calyx-lobes; stigmas greenish yellow. Fruit solitary or in pairs, on stout peduncles often nearly ½′ in length; nut nearly as broad as long, full and rounded at the ends, dark chestnut-brown, often obscurely striate, ½′—⅔′ long, inclosed for one third to one half its length in a thin cup-shaped or turbinate cup bright red-brown and lustrous on the inner surface, and covered by thin ovate light red-brown scales rounded or acute at the apex and pubescent except on their darker colored margins.

A tree, usually 50°—60° high, with a trunk rarely exceeding 3° in diameter, or rarely 100° high, with a long naked stem 3°—4° in diameter, slender tough horizontal or somewhat pendulous branches forming a narrow round-topped picturesque head, and slender branchlets dark green, lustrous, and often suffused with red when they first appear, soon glabrous, light reddish brown or light brown during their first winter and dark brown in their second year. Winter-buds ovoid, acute, about ⅛′ long, obscurely angled, and covered by closely imbricated light chestnut-brown lustrous scales erose and often ciliate on the margins. Bark on young stems and branches thin, light brown, smooth, and lustrous, becoming on old trunks ¾′—1½′ thick, and slightly divided by irregular shallow fissures into broad ridges covered by close slightly appressed light brown scales somewhat tinged with red. Wood heavy, hard, rather coarse-grained, light brown tinged with red, with thin lighter colored sapwood; occasionally used in construction, and for clapboards and shingles.

Distribution. Rich hillsides and the fertile bottom-lands of streams; Lehigh County (Allentown to Dorney’s Park), Bedford, Huntington, Franklin and Union Counties, Pennsylvania, westward through Ohio to southern Michigan, southern Wisconsin and southeastern and southern Iowa (Muscatine to Taylor County), and southward to the District of Columbia, along the Appalachian Mountains and their foothills, up to altitudes of 2200°, to the valley of the Little Tennessee River, North Carolina, and to northern Georgia (Wilkes County), and middle Tennessee; through Missouri to northeastern Kansas and southeastern Nebraska, and in northern and southern Arkansas (Fulton, Hempstead County); comparatively rare in the east; one of the most abundant Oaks of the lower Ohio basin; probably growing to its largest size in southern Indiana and Illinois.

Occasionally planted as an ornamental tree in the northern states, and hardy as far north as Massachusetts.

× Quercus Leana Nutt., scattered usually in solitary individuals from the District of Columbia and western North Carolina to southern Michigan, central and northern Illinois and southeastern Missouri, is believed to be a hybrid between this species and Quercus velutina.

× Quercus tridentata Engelm., described as a hybrid of Quercus imbricaria and Q. marilandica first found at Allenton, Saint Louis County, Missouri, occurs also near Olney, Richland County, Illinois.

× Quercus exacta Trel., believed to be a hybrid of Quercus imbricaria and Q. palustris, occurs near Olney, Richland County, Illinois, and at Crown Point, Lake County, Indiana.

21. Quercus hypoleuca Engelm.

Leaves lanceolate or oblong-lanceolate to elliptic, occasionally somewhat falcate, acute and often apiculate at apex, cuneate or rounded or cordate at the narrow base, entire or repandly serrate above the middle with occasionally small minute rigid spinose teeth, or on vigorous shoots serrate-lobed with oblique acute lobes, when they unfold light red, covered with close pale pubescence above and coated below with thick hoary tomentum, at maturity thick and firm, dark yellow-green and lustrous on the upper surface, covered on the lower with thick silvery white or fulvous tomentum, 2′—4′ long, ½′—1′ wide, with thickened revolute margins; turning yellow or brown and falling gradually during the spring after the appearance of the new leaves; petioles stout, flattened, pubescent or tomentose, ⅛′—¼′ in length. Flowers: staminate in slender aments 4′—5′ long; calyx slightly tinged with red, covered with pale hairs and divided into 4 or 5 broadly ovate rounded lobes; anthers acute, apiculate, bright red becoming yellow; pistillate mostly solitary, sessile or short-stalked, their involucral scales thin, scarious, and soft-pubescent; stigmas dark red. Fruit sessile or borne on a stout peduncle up to ½′ in length, usually solitary; nut ovoid, acute or rounded at the narrow hoary-pubescent apex, dark green and often striate when ripe, becoming light chestnut-brown in drying, ½′—⅔′ long, the shell lined with white tomentum, inclosed for about one third its length in a turbinate thick cup pubescent on the inner surface, and covered by thin broadly ovate light chestnut-brown scales rounded at apex and clothed, especially toward the base of the cup, with soft silvery pubescence.

A tree, usually 20°—30° or sometimes 60° high, with a tall trunk 10′—15′ in diameter, slender branches spreading into a narrow round-topped inversely conic head, and stout rigid branchlets coated at first with thick hoary tomentum disappearing during the first winter, becoming light red-brown often covered with a glaucous bloom and ultimately nearly black; frequently a shrub. Winter-buds ovoid, obtuse, about ⅛′ long, with thin light chestnut-brown scales. Bark ¾′—1′ thick, nearly black, deeply divided into broad ridges broken on the surface into thick plate-like scales. Wood heavy, very strong, hard, close-grained, dark brown, with thick lighter colored sapwood.

Distribution. Scattered but nowhere abundant through Pine-forests on the slopes of cañons and on high ridges usually at altitudes between 6000°—7000° above the sea on the mountains of western Texas, and of southern New Mexico and Arizona; in northern Chihuahua and Sonora.

22. Quercus agrifolia Née. Live Oak. Encina.

Leaves oval, orbicular or oblong, rounded or acute and apiculate at apex, rounded or cordate at base, entire or sinuate-dentate with slender rigid spinose teeth, when they unfold tinged with red and coated with caducous hoary tomentum, at maturity subcoriaceous, convex, dark or pale green, dull and obscurely reticulate above, paler, rather lustrous, glabrous or pubescent below, with tufts of rusty hairs in the axils of the principal veins, or sometimes covered above with fascicled hairs and coated below with thick hoary pubescence, ¾′—4′ long and ½′—3′ wide, with thickened strongly revolute margins; falling gradually during the winter and early spring; petioles stout or slender, pubescent or glabrous, ½′—1′ in length. Flowers: staminate in slender hairy aments 3′—4′ long; calyx bright purple-red in the bud, sometimes furnished with a tuft of long pale hairs at the apex, glabrous or glabrate, divided nearly to the base into 5—7 ovate acute segments reddish above the middle; pistillate sessile or short-stalked, their involucral scales bright red and covered with thick hoary tomentum, or glabrous or puberulous; stigmas bright red. Fruit sessile or nearly so, solitary or in few-fruited clusters; nut elongated, ovate, abruptly narrowed at base, gradually narrowed to the acute puberulous apex, light chestnut-brown, ¾′—1½′ long, ¼′—¾′ thick, the shell lined with a thick coat of pale tomentum, inclosed for one third its length or only at the base in a thin turbinate light brown cup coated on the inner surface with soft pale silky pubescence, and covered by thin papery scales rounded at the narrow apex, and slightly puberulous, especially toward the base of the cup.

A tree, occasionally 80°—90° high, with a short trunk 3°—4° or rarely 6°—7° in diameter, dividing a few feet above the base into numerous great limbs often resting on the ground and forming a low round-topped head frequently 150° across, and slender dark gray or brown branchlets tinged with red, coated at first with hoary tomentum persistent until the second or third year; or with a trunk, rising to the height of 30° or 40°, and crowned by a narrow head of small branches; often much smaller; frequently shrubby in habit, with slender stems only a few feet high. Winter-buds globose and usually about /16′ thick, or ovoid-oblong, acute, and sometimes on vigorous shoots nearly ¼′ in length, with thin broadly ovate closely imbricated light chestnut-brown glabrous or pubescent scales. Bark of young stems and branches thin, close, light brown or pale bluish gray, becoming on old trunks 2′—3′ thick, dark brown slightly tinged with red, and divided into broad rounded ridges separating on the surface into small closely appressed scales. Wood heavy, hard, close-grained, very brittle, light brown or reddish brown, with thick darker colored sapwood; valued and largely used for fuel.

Distribution. Usually in open groves of great extent from Sonoma County, California, southward over the coast ranges and islands to the San Pedro Mártir Mountains, Lower California; less common at the north; very abundant and of its largest size in the valleys south of San Francisco Bay and their commonest and characteristic tree; frequently covering with semiprostrate and contorted stems the sand dunes on the coast in the central part of the state; in southwestern California the largest and most generally distributed Oak-tree between the mountains and the sea, often covering low hills and ascending to altitudes of 4500° in the cañons of the San Jacinto Mountains.

Occasionally cultivated as an ornamental tree in temperate western, and in southern Europe.

23. Quercus Wislizenii A. DC. Live Oak.

Leaves narrowly lanceolate to broadly elliptic, generally oblong-lanceolate, acute or rounded and generally apiculate at apex, rounded or truncate or gradually narrowed and cuneate at base, entire, serrulate or serrate or sinuate-dentate with spreading rigid spinescent teeth, when they unfold thin, dark red, ciliate, and covered with pale scattered fascicled hairs, at maturity thick and coriaceous, glabrous and lustrous, dark green on the upper and paler and yellow-green on the lower surface, usually 1′—1½′ long and about ⅔′ wide, with obscure primary veins and conspicuous reticulate veinlets, gradually deciduous during their second summer and autumn; petioles coated at first with hoary tomentum, usually pubescent or puberulous at maturity, ⅛′ to nearly 1′ in length. Flowers: staminate in hairy aments 3′—4′ long; calyx tinged with red in the bud, divided into broadly ovate ciliate glabrous light yellow lobes shorter than the 3—6 stamens; pistillate sessile or short-stalked, their involucral scales and peduncle hoary-tomentose. Fruit sessile, short-stalked or occasionally spicate; nut slender, oblong, abruptly narrowed at base, pointed and pilose at the apex, ¾′—1½′ long, about ⅓′ thick, light chestnut-brown, often striate, the shell lined with a scanty coat of pale tomentum, more or less inclosed in the thin turbinate sometimes tubular cup ½′—1′ deep, or rarely cup-shaped and shallow, light green and puberulous within, and covered by oblong lanceolate light brown closely imbricated thin scales, sometimes toward the base of the cup thickened and rounded on the back, usually pubescent or puberulous, especially above the middle, and frequently ciliate on the margins.

A tree, usually 70°—80° high, with a short trunk 4°—6° in diameter, stout spreading branches forming a round-topped head, and slender rigid branchlets coated at first with hoary tomentum or covered with scattered fascicled hairs, puberulous or glabrous and rather light brown during their first season, gradually growing darker in their second year; usually much smaller and sometimes reduced to an intricately branched shrub, with numerous stems only a few feet tall. Winter-buds ovoid or oval, acute, ⅛′—¼′ long, with closely imbricated light chestnut-brown ciliate scales. Bark on young trees and large branches thin, generally smooth and light-colored, becoming on old trunks 2′—3′ thick, and divided into broad rounded often connected ridges separating on the surface into small thick closely appressed dark brown scales slightly tinged with red. Wood heavy, very hard, strong, close-grained, light brown tinged with red, with thick lighter colored sapwood; sometimes used for fuel.

Distribution. Lower slopes of Mt. Shasta southward through the coast region of California to the Santa Lucia Mountains, and to Santa Rosa and Santa Cruz Islands, and along the slopes of the Sierra Nevada to Kern County, up to altitudes of 2000° at the north and of 4500° at the south; as a shrub 4°—6° high with small thick leaves (var. frutescens Engelm.) on the desert slopes of the San Bernardino, San Jacinto and Cuyamaca mountains, at altitudes of 5000°—7000° above the sea, and on San Pedro Mártir in Lower California; nowhere common as a tree, but most abundant and of its largest size in the valleys of the coast region of central California at some distance from the sea, and on the foothills of the Sierra Nevada; very common as a shrub in the cañons of the desert slopes of the mountains of southern California; near the coast and on the islands small and mostly shrubby.

× Quercus morehus Kell., a supposed hybrid between Quercus Wislizenii and Q. Kelloggii occurs in Lake County, California.

24. Quercus myrtifolia Willd.

Leaves oval to oblong-obovate, acute and apiculate or broad and rounded at apex, gradually narrowed and cuneate or broad and rounded or cordate at base, entire, with much thickened revolute sometimes undulate margins, or on vigorous shoots sinuate-dentate and lobed above the middle, when they unfold, thin, dark red, coated below and on the petioles with clammy rusty tomentum and densely pubescent above, at maturity thick and coriaceous, lustrous, dark green, glabrous and conspicuously reticulate-venulose above, paler, yellow-green, or light orange-brown, glabrous or pubescent below, with tufts of rusty hairs in the axils of the veins, ½′—2′ long and ¼′—1′ wide; falling gradually during their second year; petioles stout, pubescent, yellow, rarely more than ⅛′ in length. Flowers: staminate in hoary pubescent aments 1′—1½′ long; calyx coated on the outer surface with rusty hairs and divided into 5 ovate-acute, segments shorter than the 2 or 3 stamens; pistillate sessile or nearly sessile, solitary or in pairs, their involucral scales tomentose and tinged with red. Fruit solitary or in pairs, sessile or short-stalked; nut subglobose or ovoid, acute, ¼′—½′ long, dark brown, lustrous and often striate, puberulous at apex, the shell lined with a thick coat of rusty tomentum, inclosed for one fourth to one third its length in a saucer-shaped or turbinate cup light brown and puberulous within, and covered by closely imbricated broad-ovate light brown pubescent scales ciliate on the margins and rounded at their broad apex.

A round-topped tree, rarely 40° high, with a trunk 4′—5′ or rarely up to 15′ in diameter, short or rarely long spreading branches and slender branchlets coated at first with a thick pale fulvous tomentum of articulate hairs usually persistent during the summer, light brown more or less tinged with red or dark gray, and pubescent or puberulous during their first winter, becoming darker and glabrous in their second season; more often an intricately branched shrub, with slender rigid stems 3°—4° or rarely 15°—20° high and 1′—3′ in diameter. Winter-buds ovoid or oval, gradually narrowed to the acute apex, with closely imbricated dark chestnut-brown slightly puberulous scales. Bark thin and smooth, becoming near the ground dark and slightly furrowed.

Distribution. Dry sandy ridges on the coast and islands of South Carolina to Bay Biscayne, Florida, crossing the central peninsula and from the valley of the Caloosahatchee River, westward along the coast of Florida, Alabama, and Mississippi; most abundant on the islands off the coast of east Florida, and of Alabama and Mississippi; often covering large areas with low impenetrable thickets; perhaps of its largest size in Orange County, on Jupiter Island, and on the coast west of the Apalachicola River, Florida.

25. Quercus chrysolepis Liebm. Live Oak. Maul Oak.

Leaves oblong-ovate to elliptic, acute or cuspidate at apex, cordate, rounded or cuneate at base, mostly entire on old trees, often dentate or sinuate-dentate on young trees with 1 or 2 or many spinescent teeth, the two forms often appearing together on vigorous shoots, clothed when they unfold with a thick tomentum of fulvous hairs soon deciduous from the upper and more gradually from the lower surface, at maturity thick and coriaceous, bright yellow-green and glabrous above, more or less fulvous-tomentose below during their first year, ultimately becoming glabrate and bluish white, 1′—4′ long, ½′—2′ wide, with thickened revolute margins; deciduous during their third and fourth years; petioles slender, yellow, rarely ½′ in length. Flowers: staminate in slender tomentose aments 2′—4′ long; calyx light yellow, pubescent, divided usually into 5—7 broadly ovate acute ciliate lobes often tinged with red above the middle; pistillate sessile or subsessile or rarely in short few-flowered spikes, their broadly ovate involucral scales coated with fulvous tomentum; stigmas bright red. Fruit usually solitary, sessile or short-stalked; nut ellipsoidal or ovoid, acute or rounded at the full or narrow slightly puberulous apex, light chestnut-brown, ½′—2′ long and about as thick, the shell lined with a thin coat of loose tomentum, with abortive ovules scattered irregularly over the side of the seed, inclosed only at the base in a thin hemispheric or in a thick turbinate broad-rimmed cup pale green or dark reddish brown within, and covered by small triangular closely appressed scales with a short free tip, clothed with hoary pubescence, or often hidden in a dense coat of fulvous tomentum.

A tree, usually not more than 40°—50° high, with a short trunk 3°—5° in diameter, dividing into great horizontal limbs sometimes forming a head 150° across, and slender rigid or flexible branchlets coated at first with thick fulvous tomentum, becoming during their first winter dark brown somewhat tinged with red, tomentose, pubescent, or glabrous, and ultimately light brown or ashy gray; occasionally in sheltered cañons producing trunks 8°—9° in diameter; on exposed mountain sides forming dense thickets 15°—20° high. Winter-buds broadly ovoid or oval, acute, about ⅛′ long, with closely imbricated light chestnut-brown usually puberulous scales. Bark ¾′—1½′ thick, light or dark gray-brown tinged with red, and covered by small closely appressed scales. Wood heavy, very strong, hard, tough, close-grained, light brown, with thick darker colored sapwood; used in the manufacture of agricultural implements and wagons.

Distribution. Southern Oregon, along the California coast ranges and the western slopes of the Sierra Nevada to the San Bernardino and San Jacinto mountains; of its largest size in the cañons of the coast ranges of central California and on the foothills of the Sierra Nevada; ascending to altitudes of 8000°—9000° above the sea; near the southern boundary of California, on the mountains of northern Lower California and Sonora and in Arizona (Santa Rita and Huachuca Mountains, on Beaver Creek and in Copper Cañon near Camp Verde, and in Sycamore Cañon south of Flagstaff), usually shrubby, with rigid branches, rigid coriaceous oblong or semiorbicular spinose-dentate leaves, subsessile or pedunculate fruit, with ovoid acute nuts 1′—1½′ long, their shells lined with thick or thin pale tomentum, and purple cotyledons (var. Palmeri Engelm.—Quercus Wilcoxii Rydb.)

26. Quercus tomentella Engelm.

Leaves oblong-lanceolate, acute, sometimes cuspidate or occasionally rounded at apex, broad and rounded or gradually narrowed and abruptly cuneate at base, remotely crenate-dentate with small remote spreading callous tipped teeth, or entire, when they unfold light green tinged with red, covered above with scattered pale fascicled hairs and below and on the petioles with thick hoary tomentum, at maturity thick and coriaceous, dark green, glabrous and lustrous on the upper surface, pale and covered with fascicled hairs on the lower surface, 2′—4′ long, 1′—2′ wide, with thickened strongly revolute margins, and a pubescent midrib; gradually deciduous during their third season; petioles stout, pubescent, about ½′ in length. Flowers: staminate in pubescent aments 2½′—14′ long, calyx light yellow, pubescent, divided into 5—7 ovate acute lobes; pistillate subsessile or in few-flowered spikes on short or elongated pubescent peduncles, their involucral scales like the calyx coated with fascicled hairs; stigmas red. Fruit subsessile or short-stalked; nut ovoid, broad at base, full and rounded at apex, about 1½′ long and ¾′ thick, inclosed only at the base in a cup-shaped shallow cup thickened below, light brown and pubescent on the inner surface, and covered by thin ovate acute scales, their free chestnut-brown tips more or less hidden in a thick coat of hoary tomentum.

A tree, 30°—40°, or occasionally 60° high, with a trunk 1°—2° in diameter, spreading branches forming a shapely round-topped head, and slender branchlets coated at first with hoary tomentum, becoming light brown tinged with red or orange color. Winter-buds ovoid, acute or obtuse, nearly ¼′ long, with many loosely imbricated light chestnut-brown scales more or less clothed with pale pubescence. Bark thin, reddish brown, broken into large closely appressed scales. Wood heavy, hard, close-grained, compact, pale yellow-brown, with lighter colored sapwood.

Distribution. Deep narrow cañons and high wind-swept slopes of Santa Rosa, Santa Cruz, and Santa Catalina islands, California; on Guadalupe Island off the coast of Lower California.

27. Quercus Emoryi Torr. Black Oak.

Leaves oblong-lanceolate, acute and mucronate at apex, cordate or rounded at the slightly narrowed base, entire or remotely repand-serrate with 1—5 pairs of acute rigid oblique teeth, when they unfold thin, light green more or less tinged with red and covered with silvery white tomentum, at maturity thick, rigid, coriaceous, dark green, very lustrous and glabrous or coated above with minute fascicled hairs, pale and glabrous or puberulous below, usually with 2 large tufts of white hairs at the base of the slender midrib, obscurely reticulate-venulose, 1′—2½′ long, ½′—1′ wide; falling gradually in April with the appearance of the new leaves; petioles stout, pubescent, about ¼′ in length. Flowers: staminate in hoary tomentose aments; calyx light yellow, hairy on the outer surface, divided into 5—7 ovate acute lobes; pistillate sessile or short-stalked, their involucral scales covered with hoary tomentum. Fruit ripening irregularly from June to September, sessile or short-stalked; nut oblong, oval, or ovate, narrowed at base, rounded at the narrow pilose apex, ½′—¾′ long, about ⅓′ thick, dull light green when fully grown, dark chestnut-brown or nearly black at maturity, with a thin shell lined with thick white tomentum, inclosed for from one third to one half its length in the deeply cup-shaped or nearly hemispheric cup light green and pubescent within, and covered by closely imbricated broadly ovate acute thin and scarious light brown scales clothed with short soft pale pubescence.

A tree, usually 30°—40° high, with a short trunk 2°—3° in diameter, stout rigid rather drooping branches forming a round-topped symmetrical head, and slender rigid branchlets covered at first with close hoary tomentum, bright red, pubescent or tomentose in their first winter, ultimately glabrous and dark red-brown or black; sometimes 60°—70° high, with a trunk 4°—5° in diameter, with a head occasionally 100° across; or at high altitudes or on exposed mountain slopes a low shrub. Winter-buds ellipsoidal, acute, about ¼′ long, pale pubescent toward the apex, with thin closely imbricated light chestnut-brown ciliate scales. Bark 1′—2′ thick, dark brown or nearly black, deeply divided into large oblong thick plates separating into small thin closely appressed scales. Wood heavy, strong, brittle, close-grained, dark brown or almost black, with thick bright brown sapwood tinged with red. The sweet acorns are an important article of food for Mexicans and Indians, and are sold in the towns of southern Arizona and northern Mexico.

Distribution. Mountain ranges of western Texas, southern New Mexico, Arizona south of the Colorado plateau, and of northern Mexico; in Texas common in the cañons and on the southern slopes of the Limpio and Chisos mountains; the most abundant Oak of southern New Mexico and Arizona, forming a large part of the forests covering the mountain slopes and extending from the upper limits of the mesa nearly to the highest ridges; attaining its largest size and beauty in the moist soil of sheltered cañons.

28. Quercus dumosa Nutt. Scrub Oak.

Leaves oblong, rounded and acute at apex, broad and abruptly cuneate or rounded at base, usually about ¾′ long and ½′ wide, spinescent with a few minute teeth, or undulate and entire or coarsely spinescent, with an obscure midrib and primary veins, conspicuous reticulate veinlets, and stout petioles rarely ⅛′ long; or sometimes oblong to oblong-obovate and divided by deep sinuses into 5—9 oblong acute rounded or emarginate bristle-tipped lobes, the terminal lobe 3-lobed, rounded or acute, 2′—4′ long and 1′—1½′ wide, with primary veins running to the points of the lobes, obscure reticulate veinlets, and petioles sometimes 1′ long, thin when they unfold and clothed with scattered fascicled hairs, or rarely tomentose above and coated below and on the petioles with hoary tomentum, at maturity thick and firm, dark green and glabrous on the upper surface, paler and more or less pubescent on the lower surface; mostly deciduous during the winter. Flowers: staminate in pubescent aments; calyx divided into 4—7 ovate lanceolate hairy segments; pistillate sessile or stalked, in long many-flowered tomentose spikes, their involucral scales and calyx hoary-tomentose; stigmas red. Fruit sessile or short-stalked; nut ovoid, broad at base, broad and rounded or acute at apex, ½′—1′ long, ⅓′—⅔′ thick, inclosed for one half to two thirds its length in a deep cup-shaped or hemispheric cup light brown and pubescent within, covered by ovate pointed scales coated with pale or rufous tomentum, usually much thickened, united and tuberculate, those above with free acute tips forming a fringe to the rim of the cup, or frequently with basal scales but little thickened and furnished with long free tips; in var. Alvordiana Jeps., with a nut 1½′—1⅝′ long, ¼′—½′ thick, gradually narrowed and acute at apex, inclosed only at base in a shallow cup-shaped cup.

A tree, rarely 20° high, with a trunk 12′—18′ in diameter, small branches forming a round-topped head, and slender branchlets coated at first with hoary tomentum, becoming in their first winter ashy gray or light or dark reddish brown and usually pubescent or tomentose; more often an intricately branched rigid shrub, with stout stems covered by pale gray bark and usually 6°—8° high, often forming dense thickets. Winter-buds ellipsoidal, generally acute, /16′—⅛′ long, with thin pale red often pilose and ciliate scales. Bark of the trunk bright brown and scaly.

Distribution. California; western slopes of the central Sierra Nevada; common on the coast ranges south of San Francisco Bay and on the islands off the coast of the southern part of the state, ranging inland to the borders of the Mohave Desert and to the cañons of the desert slopes of the San Bernardino and San Jacinto mountains, and southward into Lower California; arborescent only in sheltered cañons of the islands; the var. Alvordiana, in the San Emidio Cañon of the coast ranges of Kern County and on the San Carlos Range, Fresno County; north of San Francisco Bay replaced by the variety bullata Engelm. ranging to Mendocino County and to Napa valley.

× Quercus MacDonaldii Greene, a shrub or small tree with characters intermediate between those of Quercus dumosa and Q. Engelmannii, is usually considered a hybrid of these species. It occurs on Santa Cruz and Santa Catalina Islands, and in Santa Barbara, and Los Angeles Counties, California.

29. Quercus virginiana Mill. Live Oak.

Leaves oblong, elliptic or obovate, rounded or acute at apex, gradually narrowed and cuneate or rarely rounded or cordate at base, usually entire with slightly revolute margins, or rarely spinose-dentate above the middle, thin, dark green and lustrous on the upper surface, pale and pubescent on the lower surface, 2′—5′ long, ½′—2½′ wide, and inconspicuously reticulate-venulose, with a narrow yellow midrib, and few slender obscure primary veins forked and united at some distance from the margins; gradually turning yellow or brown at the end of the winter and falling with the appearance of the new leaves in the spring; petioles stout, rarely more than ¼′ in length. Flowers: staminate in hairy aments 2′—3′ long; calyx light yellow, hairy, divided into 5—7 ovate rounded segments; anthers hirsute; pistillate in spikes on slender pubescent peduncles 1′—3′ long, their involucral scales and ovate calyx-lobes coated with hoary pubescence; stigmas bright red. Fruit usually in 3—5 fruited spikes or rarely in pairs or single on stout light brown puberulous peduncles 1′—5′ long; nut ellipsoidal or slightly obovoid, narrowed at base, rounded or acute at apex, dark chestnut-brown and lustrous, about 1′ long and ⅓′ thick, inclosed for about one fourth its length in a turbinate light reddish brown cup puberulous within, its scales thin, ovate, acute, slightly keeled on the back, covered by dense lustrous hoary tomentum and ending in small closely appressed reddish tips; seed sweet, with light yellow connate cotyledons.

A tree, 40°—50° high, with a trunk 3°—4° in diameter above its swollen buttressed base, usually dividing a few feet from the ground into 3 or 4 horizontal wide-spreading limbs forming a low dense round-topped head sometimes 130° across, and slender rigid branchlets coated at first with hoary tomentum, becoming ashy gray or light brown and pubescent or puberulous during their first winter and darker and glabrous the following season; occasionally 60°—70° tall, with a trunk 6°—7° in diameter; often shrubby and occasionally not more than a foot high. Winter-buds globose or slightly obovoid, about ⅙′ long, with thin light chestnut-brown scales white and scarious on the margins. Bark of the trunk and large branches ½′—1′ thick, dark brown tinged with red, slightly furrowed, separating on the surface into small closely appressed scales. Wood very heavy, hard, strong, tough, close-grained, light brown or yellow, with thin nearly white sapwood; formerly largely and still occasionally used in shipbuilding.

Distribution. Shores of Mobjack Bay, Virginia, southward along the coast and islands to southern Florida, and along the shores of the Gulf of Mexico to northeastern Mexico, spreading inland through Texas to the valley of the Red River and to the mountains in the extreme western part of the state; on the mountains of Cuba, southern Mexico, and Central America; most abundant and of its largest size on the Atlantic and east Gulf coasts on rich hummocks and ridges a few feet above the level of the sea; abundant in Texas in the coast region, near the banks of streams, and westward toward the valley of the Rio Grande often forming the principal part of the shrubby growth on low moist soil; in sandy barren soil in the immediate vicinity of the seacoast or on the shores of salt water estuaries and bays often a shrub, sometimes bearing fruit on stems not more than a foot high (var. maritima, Sarg., and var. dentata Sarg.).

Occasionally planted as a shade and ornamental tree in the southern United States.

Variable in habit and in the size and thickness of the leaves the different forms of Quercus virginiana show little variation in their fruit. The most important of these varieties is

Quercus virginiana var. geminata Sarg.

Quercus geminata Small.

Leaves oblong-obovate to elliptic, rounded or acute at apex, cuneate or narrowed and rounded at base, occasionally slightly and irregularly dentate above the middle on vigorous shoots, conspicuously reticulate-venulose, hoary tomentose below, 1½′—3′ long, ⅓′—1′ wide, with thickened strongly revolute margins; persistent until after the leaves of the typical Q. virginiana in the same locality have all fallen; occasionally in Florida with oblong-elliptic to slightly obovate leaves 4½′—5′ long and 1′—2′ wide (f. grandifolia Sarg.). Flowers and Fruit as in the species.

A tree often 75° high with a trunk 3° in diameter, with the habit, branchlets, winter-buds and bark of the typical form; often much smaller and occasionally a shrub.

Distribution. Sandy soil; coast region of North Carolina south of the Cape Fear River, South Carolina and Georgia, and southward in Florida to Jupiter Island on the east coast and the valley of the Caloosahatchee River on the west coast; abundant and often the common Live Oak in the central part of the peninsula, at least as far south as Orange County, and westward through western Florida, southeastern and southern Alabama to the Gulf coast and islands of Mississippi.

Other varieties of Quercus virginiana are var. macrophylla Sarg., differing from the type in its much larger ovate or slightly obovate leaves rounded or acute at base, entire or occasionally repand-dentate, pale tomentose below, 3½′—4′ long and 1¼′—2½′ wide. Large trees forming groves; sandy bottoms of the Atascosa River and in flat woods above them, Pleasanton, Atascosa County, Texas: var. virescens Sarg., differing from the type in the green glabrous or rarely puberulous lower surface of the leaves and in the glabrous branchlets. A large tree in sandy soil; Gainesville, Alachua County, Sanford, Seminole County, Sumner, Levey County, Simpson’s Hummock, and near Long Key in the Everglades, Dade County, Florida: var. eximea Sarg., differing from the type in its narrow elliptic to narrow oblong-obovate leaves and pale bark; a tree rarely 20° high, with a trunk 8′—12′ in diameter; rarely a shrub; dry sandy open woods, near Springfield, Livingston Parish and near Hammond, Tangipahoa Parish, eastern Louisiana. The following small shrubby small-leaved forms are recognized: var. fusiformis Sarg., with oblong-ovate leaves acute at apex, rounded or cuneate at base, entire or occasionally dentate, and pale pubescent below, and small fruit; dry limestone ridges and flat-topped hills of the Edwards Plateau (Kerr and Comal Counties), western Texas: var. dentata Chapm., distinct in the oblong-obovate repand-dentate lower leaves with large triangular teeth, acute at the broad apex, often 4′ long and 1¼′ wide at the base of the stems, and much larger than the oblong-lanceolate entire upper leaves; common in sterile pine-barrens near the coast of Florida: var. maritima Sarg., with oblong-obovate or rarely lanceolate leaves, acute and apiculate or rounded at apex, cuneate at base, and entire or slightly and irregularly toothed above the middle; fruit solitary or in pairs, or rarely in elongated spikes (Quercus succulenta Small); sandy barrens near the coast, South Carolina to Miami, Dade County, Florida: var. pygmaea Sarg., with oblong-obovate leaves, cuneate at base, 3—5 lobed at apex with small acute lobes, or rarely elliptic and entire, and nearly sessile fruit, the nut inclosed nearly to the apex; a shrub rarely 3° high; Pine-woods in sandy soil; widely distributed in Florida.

30. Quercus reticulata H. B. K.

Leaves broadly obovate, obtuse and rounded or rarely acute at apex, usually cordate or occasionally rounded at the narrow base, repandly spinose-dentate above the middle or only toward the apex with slender teeth, and entire below, when they unfold coated with dense fulvous tomentum, at maturity thick, firm, and rigid, dark blue and covered with scattered fascicled hairs above, paler and coated with thick fulvous pubescence below, 1′—5′ long, ¾′—4′ broad, with a thick midrib, and primary veins running to the points of the teeth or arcuate and united within the slightly revolute margins, and very conspicuous reticulate veinlets; petioles stout about ¼′ in length. Flowers: staminate in short tomentose aments in the axils of leaves of the year; calyx light yellow, hirsute, with pale hairs, divided into 5—7 ovate acute segments; pistillate in spikes on elongated peduncles, clothed like their involucral scales with hoary tomentum; stigmas dark red. Fruit usually in many-fruited spikes or occasionally in pairs or rarely solitary, on slender hirsute or glabrous peduncles 2′—5′ long; nut oblong, rounded or acute at the pilose apex, broad at base, about ½′ long, inclosed for about one fourth its length in a shallow cup-shaped cup dark brown and pubescent within, hoary tomentose without and covered by small ovate acute scales, with thin free scarious tips, slightly thickened and rounded on the back at the bottom of the cup.

A tree, rarely more than 40° high, with a trunk 1° in diameter, and stout branchlets coated at first with thick fulvous tomentum, light orange color and more or less thickly clothed with pubescence during their first winter, becoming ashy gray or light brown; in the United States usually shrubby in habit and sometimes only a few feet tall; becoming on the Sierra Madre of Mexico a large tree. Winter-buds ovoid to oval, often surrounded by the persistent stipules of the upper leaves, about ⅛′ long, with thin loosely imbricated light red scales ciliate on the margins. Bark about ¼′ thick, dark or light brown, and covered by small thin closely appressed scales. Wood very heavy, hard, close-grained, dark brown, with thick lighter colored sapwood.

Distribution. Near the summits of the mountain ranges of southeastern New Mexico (Mogollon Mountains) and southeastern Arizona, and southward in Mexico.

31. Quercus Toumeyi Sarg.

Leaves ovate or ovate-oblong or oval, acute and apiculate at apex, rounded or cordate at base, entire with thickened slightly revolute margins, or remotely spinulose-dentate, often minutely 3-toothed at apex, thin but firm in texture, light blue-green, glabrous and lustrous above, pale and puberulous below, conspicuously reticulate-venulose; ½′—¾′ long, ¼′—½′ wide; falling early in spring with the appearance of the new leaves; petioles stout, tomentose, about /16′ in length. Flowers unknown. Fruit sessile, solitary or in pairs, ripening in June; nut oval or ovoid, ½′—⅔′ long, ¼′ thick, light brown and lustrous, furnished at the acute apex with a narrow ring of pale pubescence, inclosed for about one half its length in a thin shallow tomentose cup light green and pubescent within, and covered by thin ovate regularly and closely imbricated light red-brown scales ending in a short rounded tip and coated on the back with pale tomentum.

A tree, 25°—30° high, with a short trunk 6′—8′ in diameter, dividing not far from the ground into numerous stout wide-spreading branches forming a broad irregular head, and slender branchlets bright red-brown more or less thickly coated with pale tomentum at midsummer, covered during their second and third years with thin dark brown nearly black bark broken into small thin closely appressed scales. Wood light brown, with thick pale sapwood.

Distribution. Forming an open forest on the Mule Mountains, Cochise County, southeastern Arizona.

32. Quercus arizonica Sarg. White Oak.

Leaves oblong-lanceolate to broadly obovate, generally acute or sometimes rounded at apex, rounded or cordate at base, repandly spinose-dentate usually, except on vigorous shoots, only above the middle or toward the apex, or entire and sometimes undulate on the margins, when they unfold light red clothed with bright fulvous tomentum and furnished with dark dental glands, at maturity thick, firm and rigid, dull dark blue-green and glabrate above, duller and covered with thick fulvous or pale pubescence below, 1′—4′ long, ½′—2′ wide, with a broad yellow midrib, slender primary veins, arcuate and united near the thickened revolute margins, and coarsely reticulate veinlets; falling in the early spring just before the appearance of the new leaves; petioles stout, tomentose, ¼′—½′ in length. Flowers: staminate in tomentose aments 2′—3′ long; calyx pale yellow, pubescent, and divided into 4—7 broad acute ciliate lobes; anthers red or yellow; pistillate on short stems tomentose like their involucral scales. Fruit sessile or on hoary-tomentose stems rarely ½′ long, usually solitary, ripening irregularly from September to November; nut oblong, oval or slightly obovoid, obtuse and rounded at the puberulous apex, ¾′—1′ long, ½′ thick, dark chestnut-brown, lustrous and often striate, soon becoming light brown, inclosed for half its length in a cup-shaped or hemispheric cup light brown and pubescent within, covered by regularly and closely imbricated scales coated with pale tomentum and ending in thin light red pointed tips, those below the middle of the cup much thickened and rounded on the back; seed dark purple, very astringent.

A tree, occasionally 50°—60° tall, with a trunk 3°—4° in diameter, and thick contorted branches spreading nearly at right angles and forming a handsome round-topped symmetrical head, and stout branchlets clothed at first with thick fulvous tomentum persistent during their first winter, reddish brown or light orange color and pubescent or puberulous in their second season, ultimately glabrous and darker; usually not more than 30°—40° tall; at high elevations reduced to a low shrub. Winter-buds subglobose, about /16′ long, with loosely imbricated bright chestnut-brown puberulous scales ciliate on the margins. Bark of young stems and branches thin, pale, scaly with small appressed scales, becoming on old trunks about 1′ thick and deeply divided by narrow fissures into broad ridges broken into long thick plate-like scales pale or ashy gray on the surface. Wood heavy, strong, hard, close-grained, dark brown or nearly black, with thick lighter colored sapwood; used only for fuel.

Distribution. The most common and generally distributed White Oak of southern New Mexico and Arizona, covering the slopes of cañons of mountain ranges at altitudes of from 5000°—10,000° above the sea, often ascending nearly to the summits of the high peaks; and in northern Mexico.

33. Quercus oblongifolia Torr. White Oak.

Leaves ovate, elliptic, or slightly obovate, rounded and occasionally emarginate or acute at apex, usually cordate or occasionally rounded at base, entire and sometimes undulate with thickened revolute margins, or remotely dentate with small callous teeth, on vigorous shoots and young plants oblong, rounded or cuneate at the narrow base, coarsely sinuate or undulate-toothed or 3-toothed at the broad apex and entire below, when they unfold bright red and coated with deciduous hoary tomentum, at maturity thin and firm, blue-green and lustrous above, paler below, 1′—2′ long, ½′—¾′ wide, or on vigorous shoots sometimes 3′—4′ long, with a prominent pale midrib, slender primary veins, and conspicuous reticulate veinlets; persistent during the winter without change of color, gradually turning yellow in the spring and falling at the appearance of the new leaves; petioles stout, nearly terete, about ¼′ in length. Flowers: staminate in short hoary-tomentose aments; calyx bright yellow, pilose, divided into 5 or 6 laciniately cut or entire acute segments tinged with red above the middle; pistillate usually sessile, or on peduncles tomentose like the involucral scales; stigmas bright red. Fruit usually solitary and sessile, rarely long-stalked; nut ovoid, ellipsoidal, or slightly obovoid, full and rounded at apex surrounded by a narrow ring of white pubescence, dark chestnut-brown, striate, and very lustrous, soon becoming light brown in drying, ½′—¾′ long, about ⅓′ thick, inclosed for about one third its length in a cup-shaped or rarely turbinate thin cup yellow-green and pubescent on the inner surface and covered by ovate-oblong scales slightly thickened on the back, coated with hoary tomentum and ending in thin acute bright red tips ciliate on the margins and sometimes forming a minute fringe to the rim of the cup.

A tree, rarely more than 30° high, with a short trunk 18′—20′ in diameter, many stout spreading often contorted branches forming a handsome round-topped symmetrical head, and slender rigid branchlets coated at first with pale or fulvous tomentum, light red-brown, dark brown or dark orange color in their first winter, becoming ashy gray in their second or third year. Winter-buds subglobose, /16′—⅛′ long, with thin light chestnut-brown scales. Bark ¾′—1¼′ thick, ashy gray, and broken into small nearly square or oblong close plate-like scales. Wood very heavy, hard, strong, brittle, dark brown or nearly black, with thick brown sapwood; sometimes used as fuel.

Distribution. Chisos Mountains, western Texas, southeastern New Mexico, southern Arizona, and southward into northern Mexico; comparatively rare in Texas; abundant on the foothills of the mountain ranges of southern New Mexico and Arizona at altitudes of about 5000°, and dotting the upper slopes of the mesa where narrow cañons open to the plain.

34. Quercus Engelmannii Greene. Evergreen Oak.

Leaves oblong to obovate, usually obtuse and rounded or sometimes acute at apex, gradually or abruptly cuneate or rounded or cordate at base, entire, often undulate, or sinuate-toothed with occasionally rigid teeth, or at the ends of sterile branches frequently coarsely crenately serrate with incurved teeth, or rarely lobed with acute oblique rounded lobes, when they unfold bright red and coated with thick pale rufous tomentum, at maturity thick, dark blue-green and glabrous or covered with fascicled hairs above, pale, usually yellow-green and clothed with light brown pubescence, or puberulous or often glabrous below, 1′—3′ long, ½′—2′ wide; deciduous in the spring with the appearance of the new leaves; petioles slender, tomentose, becoming pubescent, ¼′—½′ in length. Flowers: staminate in slender hairy aments 2′—3′ long; calyx light yellow, pilose, with lanceolate acute segments; pistillate on slender peduncles, clothed like their involucral scales with dense pale tomentum. Fruit sessile or on slender pubescent peduncles sometimes ¾′ long; nut oblong, gradually narrowed and acute or broad rounded and obtuse at apex, broad or narrow at base, dark chestnut-brown more or less conspicuously marked by darker longitudinal stripes, turning light chestnut-brown in drying, ¾′—1′ long, about ½′ thick, inclosed for about half its length in a deep saucer-shaped, cup-shaped or turbinate cup light brown and puberulous within, and covered by ovate light brown scales coated with pale tomentum, usually thickened, united and tuberculate at the base of the cup, and near its rim produced into small acute ciliate tips.

A tree, 50°—60° high, with a trunk 2°—3° in diameter, thick branches spreading nearly at right angles and forming a broad rather irregular head, and stout rigid branchlets coated at first with hoary tomentum, light or dark brown tinged with red and pubescent during their first winter, becoming glabrous and light brown or gray in their second or third year. Winter-buds oval or ovoid, about ⅛′ long, with thin light red pubescent scales. Bark 1½—2′ thick, light gray tinged with brown, deeply divided by narrow fissures and separating on the surface into small thin appressed scales. Wood very heavy, hard, strong, close-grained, brittle, dark brown or nearly black, with thick lighter brown sapwood; used only for fuel.

Distribution. Low hills of southwestern California west of the coast range, occupying with Quercus agrifolia Née, a belt about fifty miles wide, and extending to within fifteen or twenty miles of the coast, from the neighborhood of Sierra Madre and San Gabriel, Los Angeles County, to the mesa east of San Diego; in northern Lower California.

35. Quercus Douglasii Hook. & Arn. Blue Oak. Mountain White Oak.

Leaves oblong, acute or rounded at apex, gradually narrowed and cuneate or broad and rounded or subcordate at base, divided by deep or shallow, wide or narrow sinuses acute or rounded in the bottom into 4 or 5 broad or narrow acute or rounded often mucronate lobes, 2′—5′ long, 1′—1¾′ wide, or oval, oblong or obovate, rounded or acute at apex, equally or unequally cuneate or rounded at base, regularly or irregularly sinuate-toothed with rounded acute rigid spinescent teeth, or denticulate toward the apex, 1′—2′ long, ¼′—1′ wide, when they unfold covered by soft pale pubescence, at maturity thin, firm and rather rigid, pale blue, with scattered fascicled hairs above, often yellow-green and covered by short pubescence below, with a hirsute or puberulous prominent midrib and more or less conspicuous reticulate veinlets; petioles stout, tomentose, ¼′—½′ in length. Flowers: staminate in hairy aments 1½′—2′ long; calyx yellow-green, coated on the outer surface with pale hairs, deeply divided into broad acute laciniately cut segments; pistillate in short few-flowered spikes coated like the involucral scales with hoary tomentum. Fruit sessile or short-stalked, solitary or in pairs; nut ellipsoidal, sometimes ventricose, with a narrow base, gradually narrowed and acute at apex, ¾′—1′ long, ½′—1′ thick, or often ovoid and acute, green and lustrous, turning dark chestnut-brown in drying, with a narrow ring of hoary pubescence at apex, inclosed only at base in a thin shallow cup-shaped cup light green and pubescent on the inner surface, covered on the outer by small acute and usually thin or sometimes, especially in the south, thicker tumid scales coated with pale pubescence or tomentum and ending in thin reddish brown tips.

A tree, usually 50°—60°, rarely 80°—90° high, with a trunk 3°—4° in diameter, short stout branches spreading nearly at right angles and forming a dense round-topped symmetrical head, stout branchlets brittle at the joints, coated at first with short dense hoary tomentum, dark gray or reddish brown and tomentose, pubescent, or puberulous during their first winter, becoming ultimately ashy gray or dark brown; frequently not more than 20°—30° high, and sometimes, especially southward shrubby, in habit. Winter-buds ovoid, obtuse, ⅛′—¼′ long, with light rather bright red pubescent scales. Bark ½′—1′ thick, generally pale, and covered by small scales sometimes tinged with brown or light red. Wood hard, heavy, strong, brittle, dark brown, becoming nearly black with exposure, with thick light brown sapwood; largely used as fuel.

Distribution. Scattered over low hills, dry mountain slopes and valleys; California, Mendocino County, and the upper valley of the Sacramento River, southward along the western slopes of the Sierra Nevada up to elevations of 4000°, and through valleys of the coast ranges to the Tehachapi Pass, the borders of the Mohave Desert (Sierra de la Liebre) and the neighborhood of San Fernando, Los Angeles County; most abundant and of its largest size in the valleys between the coast mountains and the interior ridges of the coast ranges south of the Bay of San Francisco.

× Quercus jolonensis Sarg. with characters intermediate between those of Quercus Douglasii and Quercus lobata and believed to be a hybrid of those species occurs, with a number of large trees, at Jolon and between Jolon and King City, Monterey County, California.

36. Quercus Vaseyana Buckl. Shin Oak.

Quercus undulata var. Vaseyana Rydb.

Leaves oblong, rarely oblong-obovate, acute or rounded at apex, cuneate at base, undulately lobed with small acute lobes pointing forward, rarely nearly entire, when they unfold covered above with short fascicled hairs sometimes persistent until midsummer, and tomentose below, and at maturity thin, pale gray-green, glabrous and lustrous above, pale pubescent below, 1′—1½′ long and ½′—¾′ wide; deciduous late in winter or in early spring; petioles covered with fascicled hairs when they first appear, becoming glabrous, ¼′ in length. Flowers: staminate in villose aments 1′—1¼′ long; calyx deeply divided into 4 or 5 ovate scarious lobes rounded at apex and shorter than the stamens; pistillate on short tomentose peduncles, their involucral scales ovate, acute, pubescent, shorter than the calyx-lobes; stigmas red. Fruit solitary or in pairs, sessile or short-stalked; nut ellipsoidal and only slightly narrowed at the rounded ends to oblong and slightly ovoid or obovoid, ½′—¾′ in length, ¼′—½′ in diameter, pale chestnut-brown and lustrous, the base only inclosed in the thin, saucer-shaped to cup-shaped cup, puberulous on the inner surface, covered with closely appressed ovate acute hoary tomentose scales, on some individuals abruptly contracted into short acute red-brown nearly glabrous tips.

A tree, rarely 15°—20° high, usually a shrub only 1°—3° tall, spreading into great thickets, with slender branchlets thickly covered with matted fascicled hairs during their first season, and light gray and glabrous or puberulous in their second year. Winter-buds ovoid or obovoid, about ⅛′ long, with red-brown scales ciliate on the margins. Bark rough, deeply furrowed and scaly.

Distribution. Limestone slopes and ridges or in sheltered cañons; western Texas; Kimble, Real, Kendall, Kerr, Uvalde, Edwards, Menard and Valverde Counties.

37. Quercus Mohriana Rydb. Shin Oak.

Leaves oblong-obovate to elliptic or lanceolate, acute, acuminate or rounded at apex, rounded or cuneate and often unsymmetrical at base, entire, undulate, sinuately toothed with triangular apiculate teeth, or occasionally irregularly lobed above the middle with rounded lobes, thick, gray-green, lustrous and covered above with short fascicled hairs, and densely hoary tomentose below, 2°—4° long, ½′—1′ wide, with a stout midrib thickly covered with fascicled hairs, sometimes becoming glabrous, slender primary veins and reticulate veinlets; petioles stout, hoary tomentose, ⅛′—¼′ in length. Flowers: staminate in short hoary tomentose aments; calyx densely villose, deeply divided into broad ovate lobes rounded at apex; anthers red; pistillate on hoary tomentose peduncles, with hairy bracts and calyx-lobes. Fruit solitary or in pairs, nearly sessile or raised on a pubescent peduncle ½′—¾′ in length; nut ellipsoidal or ovoid, broad and rounded at the ends, light chestnut-brown, lustrous, ⅓′—½′ long, ¼′—⅓′ thick, inclosed for from half to two thirds its length in the hemispheric to cup-shaped cup, hoary tomentose on the inner surface, and covered with small closely appressed acute hoary tomentose scales much thickened below the middle of the cup, thin and much smaller toward its rim.

A tree, rarely 18°—20° high, with a trunk rarely 1° in diameter, small spreading and ascending branches forming a round-topped head, and slender branchlets thickly coated during their first season with fascicled hairs, dark gray-brown and pubescent in their second season and ultimately gray and glabrous; usually a low shrub spreading into thickets. Winter-buds broad-ovoid, obtuse, pale pubescent. Bark thin, pale, rough, deeply furrowed.

Distribution. On dry limestone hills, usually not more than 18° high with spreading branches; on deep sand, often not more than 3° high with more erect stems, often covering thousands of acres; only a tree in the protection of ledges in deep ravines and on steep hillsides; northwestern Texas (Tom Green, Coke, Nolan, Howard, Armstrong, and Wheeler Counties); central Texas (Bryan, Brazos County); southwestern Oklahoma (Beckham County).

38. Quercus Laceyi Small.

Leaves oblong to oblong-obovate, usually with two pairs of small rounded lateral lobes, occasionally 3-lobed toward the apex, rarely nearly entire, narrowed and rounded at apex, rounded, cuneate or rarely cordate at the gradually narrowed base, coated below when they unfold with loose white tomentum, soon glabrous, at maturity thin, blue-green above, yellow-green below, 2′—3′ long, ¾′—2′ wide, with a slender midrib and primary veins, and conspicuous reticulate veinlets; deciduous late in the autumn; on vigorous shoots sometimes 6′—7′ long and 3′—4′ wide; petioles glabrous or sparingly villose, ¼′—⅓′ in length. Flowers: staminate in slightly villose aments 2′—2½′ long; calyx deeply divided into 4 or 5 ovate acuminate lobes shorter than the stamens; pistillate flowers not seen. Fruit solitary or in pairs, sessile or raised on a stem up to ½′ in length; nut ellipsoidal or oblong-ovoid, rounded at apex, slightly narrowed and nearly truncate at base, light chestnut-brown and lustrous, ¾′—1′ long, ⅓′—½′ in diameter, the base inclosed in the thick, cup-shaped to rarely saucer-shaped cup, tomentose on the inner surface, covered with acute much thickened pale tomentose scales.

A tree, 30°—45° high, with a trunk 20′—30′ in diameter, heavy erect and spreading branches and slender branchlets villose when they first appear, soon becoming glabrous and red-brown or gray during their second season; often a tall shrub with numerous stems. Winter-buds ovoid, acute, ⅙′ long, with chestnut-brown scales ciliate on the margins. Bark gray, thick, deeply ridged or checkered.

Distribution. Rocky banks of streams, the steep sides of cañons and on limestone bluffs; common in the southern and southwestern parts of the Edwards Plateau, western Texas (Kendall, Kerr, Bandera, Uvalde, Menard, Kemble, Real and Edwards Counties); easily distinguished in the field by the peculiar smoky or waxy appearance of the foliage.

39. Quercus annulata Buckl.

Quercus breviloba Sarg.

Leaves oblong to oblong-obovate or elliptic, rounded or acute at apex, cuneate or rounded at base, entire, undulate, slightly lobed with rounded or acute lobes, or 3-lobed, when they unfold covered above with fascicled hairs and tomentose below, and at maturity green, glabrous and lustrous above, green and pubescent below on lower branches, often pale or hoary tomentose on upper branches, 1¼′—2½′ long, ½′—1¼′ wide; petioles covered when they first appear with fascicled hairs, soon glabrous, ¼′—½′ in length; on vigorous branchlets sometimes thinner, glabrous, divided into broad rounded lateral lobes, gradually narrowed and cuneate at the long base, 4′ long and 2½′ wide. Flowers: staminate in pubescent aments 1′—2′ long; calyx deeply divided in villose rounded lobes, shorter than the stamens; anthers red; pistillate on tomentose peduncles, their scales rounded, tomentose; stigmas red. Fruit solitary or in 2 or 3-fruited clusters, sessile or short-stalked, oblong-ovoid to ellipsoidal, slightly narrowed and rounded at apex, light yellow-brown and lustrous, ¾′—1′ long, ⅓′—½′ in diameter; inclosed for about a quarter of its length in the cup-shaped cup, tomentose on the inner surface, covered with acute tomentose scales somewhat thickened and closely appressed below the middle of the cup, their tips chestnut-brown, free and often glabrous.

A tree, 20°—30° tall with a trunk rarely more than 1° in diameter, small spreading often slightly pendulous branches forming a round-topped head, and slender branches covered when they first appear with fascicled hairs, soon becoming glabrous and gray or grayish brown; the large stems often surrounded by a ring of smaller stems produced from its roots; more often a shrub than a tree spreading into broad thickets. Winter-buds ovoid to ellipsoidal, acute, ⅛′—¼′ long, with closely imbricated chestnut-brown puberulous scales ciliate on the margins. Bark thick, rough, deeply ridged.

Distribution. Dry limestone hills and bluffs; central and western Texas, from the neighborhood of Dallas, Dallas County, and Palo Pinto County to Kendall, Kerr, Brown, Bandera, Real and Menard Counties.

40. Quercus Durandii Buckl.

Quercus breviloba Sarg. in part.

Leaves thin, obovate to elliptic, entire, 3-lobed toward the rounded or acute apex or irregularly laterally lobed, the three forms appearing on different branches of the same tree, on lower branches usually lobed, dark green and lustrous above, often green and glabrous below, sometimes 6′ or 7′ long and 3′ or 3½′ wide, on upper branches mostly entire, white and pubescent or tomentose below, 2½′—3′ long, ½′—1½′ wide; falling late in the autumn; petioles glabrous, ⅓′—¼′ in length. Flowers: staminate in slender villose aments 3′—4′ in length; calyx deeply divided into acute villose lobes shorter than the stamens; pistillate on a short tomentose peduncle, the linear acuminate bract and involucral scales hoary-tomentose; stigmas red. Fruit solitary or in pairs, short-stalked or nearly sessile; nut ovoid, or slightly obovoid, rounded or rarely acute at apex, nearly truncate at base, pale chestnut-brown, lustrous, ½′—⅔′ long, ⅓′—½′ thick, barely inclosed at base in the thin, shallow saucer-shaped cup, pale tomentose on the inner surface, and covered with small acuminate closely appressed tomentose scales slightly thickened on the back.

A tree, often 60°—90° high with a tall trunk 2°—3° in diameter, comparatively small branches, the lower horizontal, the upper ascending, forming a dense round-topped handsome head, and slender pale gray-brown branchlets covered when they first appear with fascicled hairs, soon glabrous, or puberulous during their first season, and darker in their second season. Winter-buds ovoid, acute, ¼′—⅓′ long with dark chestnut-brown rounded scales ciliate on the margins. Bark thin, light gray or nearly white and broken into thin loosely appressed scales.

Distribution. East of the Mississippi River scattered on rich limestone prairies; westward on the well drained soil of river bottoms, and often on low hummocks; near Augusta, Richmond County, and De Soto, Sampson County, Georgia; West Point, Clay County, Columbus, Muscogee County, Brookville, Noxubesco County, and near Natchez, Adams County, Mississippi; McNab, Hempstead County, Arkansas; Natchitoches, Natchitoches Parish, western Louisiana; coast region of eastern Texas to the bottoms of the Guadalupe River (Victoria, Victoria County), ranging inland to San Saba County and to the neighborhood of Dallas, Dallas County; on the mountains near Monterey, Nuovo Leon; rare and local.

41. Quercus Chapmanii Sarg.

Leaves oblong to oblong-obovate, rounded at the narrow apex, narrowed and cuneate or rounded or broad and rounded at base, entire with slightly undulate margins, or obscurely sinuate-lobed above the middle, when they unfold coated below with thick bright yellow pubescence and covered above with pale fascicled deciduous hairs, at maturity thick and firm or subcoriaceous, dark green, glabrous and lustrous above, light green or silvery white and glabrous below except on the slender often pubescent midrib, usually 2′—3′ long and 1′ wide, but varying from 1′—3′ in length and ¾′—1′ in width; falling gradually during the winter or sometimes persistent until the appearance of the new leaves in the spring; petioles tomentose, rarely ⅛′ in length. Flowers: staminate in short hirsute aments; calyx hirsute, divided into 5 acute laciniately cut segments; anthers hirsute; pistillate sessile or short-stalked, their involucral scales coated with dense pale tomentum. Fruit usually sessile, solitary or in pairs; nut oval, about ⅝′ long and ⅜′ thick, pubescent from the obtuse rounded apex nearly to the middle, inclosed for nearly half its length in the deep cup-shaped light brown cup slightly pubescent on the inner surface, and covered by ovate-oblong pointed scales thickened on the back, especially toward the base of the cup, and coated with pale tomentum except on their thin reddish brown margins.

Occasionally a tree, 50° high, with a trunk 1° in diameter, stout branches forming a round-topped head, and slender branchlets coated at first with dense bright yellow pubescence, becoming light or dark red-brown and puberulous during their first winter and ultimately ashy gray; more often a rigid shrub sometimes only 1°—2° tall. Winter-buds ovoid, acute, obtuse, about ⅛′ long, with glabrous or puberulous light chestnut-brown scales. Bark dark or pale, separating freely into large irregular plate-like scales.

Distribution. Sandy barrens usually in the neighborhood of the coast; Bluffton, Beaufort County, South Carolina, Colonels Islands, Liberty County, Georgia, southward along the east coast of Florida to the shores of Indian River; on the west coast from the valley of the Caloosahatchee River to the shores of Pensacola Bay, and in the interior of the peninsular from Lake County to De Soto County (neighborhood of Sebring); rare and local on the Atlantic coast; comparatively rare in the interior of the Florida peninsular; abundant in western Florida from the shores of Tampa Bay to those of Saint Andrews Bay.

42. Quercus macrocarpa Michx. Burr Oak. Mossy Cup Oak.

Leaves obovate or oblong, cuneate or occasionally narrow and rounded at base, divided by wide sinuses sometimes penetrating nearly to the midrib into 5—7 lobes, the terminal lobe large, oval or obovate, regularly crenately lobed, or smaller and 3-lobed at the rounded or acute apex, when they unfold yellow-green and pilose above and silvery white and coated below with long pale hairs, at maturity thick and firm, dark green, lustrous and glabrous, or occasionally pilose on the upper surface, pale green or silvery white and covered on the lower surface with soft pale or rarely rufous pubescence, 6′—12′ long, 3′—6′ wide, with a stout pale midrib sometimes pilose on the upper side and pubescent on the lower, large primary veins running to the points of the lobes, and conspicuous reticulate veinlets; turning dull yellow or yellowish brown in the autumn; petioles stout, ⅓′—1′ in length. Flowers: staminate in slender aments 4′—6′ long, their yellow-green peduncles coated with loosely matted pale hairs; calyx yellow-green, pubescent, deeply divided into 4—6 acute segments ending in tufts of long pale hairs; pistillate sessile or stalked, their involucral scales broadly ovate, often somewhat tinged with red toward the margins and coated, like the peduncles, with thick pale tomentum; stigmas bright red. Fruit usually solitary, sessile or long-stalked, exceedingly variable in size and shape; nut ellipsoidal or broad-ovoid, broad at the base and rounded at the obtuse or depressed apex covered by soft pale pubescence, ⅗′ long and ⅓′ thick at the north, sometimes 2′ long and 1½′ thick in the south, its cup thick or thin, light brown and pubescent on the inner surface, hoary-tomentose and covered on the outer surface by large irregularly imbricated ovate pointed scales, at the base of the cup thin and free or sometimes much thickened and tuberculate, and near its rim generally developed into long slender pale awns forming on northern trees a short inconspicuous and at the south a long conspicuous matted fringe-like border, inclosing only the base or nearly the entire nut.

A tree, sometimes 170° high, with a trunk 6°—7° in diameter, clear of limbs for 70°—80° above the ground, a broad head of great spreading branches, and stout branchlets coated at first with thick soft pale deciduous pubescence, light orange color, usually glabrous or occasionally puberulous during their first winter, becoming ashy gray or light brown and ultimately dark brown, sometimes developing corky wings often 1′—1½′ wide; usually not more than 80° high, with a trunk 3°—4° in diameter; toward the northwestern limits of its range sometimes a low shrub. Winter-buds broadly ovoid, acute or obtuse, ⅛′—¼′ long, with light red-brown scales coated with soft pale pubescence. Bark 1′—2′ thick, deeply furrowed and broken on the surface into irregular plate-like brown scales often slightly tinged with red. Wood heavy, strong, hard, tough, close-grained, very durable, dark or rich light brown, with thin much lighter colored sapwood; used in ship and boat-building, for construction of all sorts, cabinet-making, cooperage, the manufacture of carriages, agricultural implements, baskets, railway-ties, fencing, and fuel.

Distribution. Low rich bottom-lands and intervales, or rarely in the northwest on low dry hills; Nova Scotia and New Brunswick southward to the valley of the Penobscot River, Maine, the shore of Lake Champlain, Vermont, western Massachusetts, central, southern and western Pennsylvania, northern Delaware, northern West Virginia (Hardy and Grant Counties), prairies of Caswell County, North Carolina, and middle Tennessee, and westward through the valley of the Saint Lawrence River and along the northern shores of Lake Huron to southern Manitoba, through western New York and Ohio, northern Michigan, to Minnesota (except in the northeastern counties), eastern and northwestern Nebraska, the Black Hills of South Dakota, the Turtle Mountains of North Dakota, and northeastern Wyoming, and to central Kansas, the valley of the north Fork of the Canadian River (Canton, Blaine County, and Seiling, Dewey County), Oklahoma, and the valley of the San Saba River, (Menard County and Callahan County), Texas; attaining its largest size in southern Indiana and Illinois; the common Oak of the “oak openings” of western Minnesota, and in all the basin of the Red River of the North, ranging farther to the northwest than the other Oaks of eastern America; common and generally distributed in eastern Nebraska, and of a large size in cañons or on river bottoms in the extreme northwestern part of the state; the most generally distributed Oak in southern Wisconsin, and in Kansas growing to a large size in all the eastern part of the state.

Occasionally planted as an ornamental tree in the eastern United States and in South Africa.

× Quercus Andrewsii Sarg., believed to be a hybrid of Quercus macrocarpa and Q. undulata Torr., in habit and characters intermediate between those of its supposed parents with which it grows, occurs at Seiling, Dewey County, western Oklahoma.

× Quercus guadalupensis Sarg., with characters intermediate between those of Quercus macrocarpa and Q. stellata and evidently a hybrid of these species, occurs at Fredericksburg Junction in the valley of the Guadalupe River, Kendall County, Texas.

× Quercus Hillii Trel., believed to be a hybrid of Quercus macrocarpa and Q. Muehlenbergii, has been found at Roby, Lake County, Indiana, and near Independence, Jackson County, Missouri.

43. Quercus lyrata Walt. Overcup Oak. Swamp White Oak.

Leaves oblong-obovate, gradually narrowed and cuneate at base, divided into spreading or ascending lobes by deep or shallow sinuses rounded, straight, or oblique on the bottom, the terminal lobe oblong-ovate, usually broad, acute or acuminate at the elongated apex, and furnished with 2 small entire nearly triangular lateral lobes, the upper lateral lobes broad, more or less emarginate, or acuminate and entire or slightly lobed and much longer than the acute or rounded lower lobes, when they unfold bronze-green and pilose above with caducous hairs, and coated below with thick pale tomentum, at maturity thin and firm, dark green and glabrous above, silvery white and thickly coated with pale pubescence, or green and often nearly glabrous below, 7′—10′ long, 1′—4′ wide; turning yellow or scarlet and orange in the autumn; petioles glabrous or pubescent, ⅓′—1′ in length. Flowers: staminate in slender hairy aments 4′—6′ long; calyx light yellow, coated on the outer surface with pale hairs and divided into acute segments; pistillate sessile or stalked, their involucral scales covered, like the peduncles, with thick pale tomentum. Fruit sessile or borne on slender pubescent peduncles sometimes 1½′ in length; nut subglobose to ovoid or rarely to ovoid-oblong, ½′—1′ long, usually broader at base than long, light chestnut-brown, more or less covered above the middle with short pale pubescence, entirely or for two thirds of its length inclosed in the ovoid, nearly spherical or deep cup-shaped thin cup, bright red-brown and pubescent on the inner surface, hoary-tomentose and covered on the outer by ovate united scales produced into acute tips, much thickened and contorted at its base, gradually growing thinner and forming a ragged edge to the thin often irregularly split rim of the cup.

A tree, rarely 100° high, with a trunk 2°—3° in diameter, generally divided 15°—20° above the ground into comparatively small often pendulous branches forming a handsome symmetrical round-topped head, and slender branchlets green more or less tinged with red and pilose or pubescent when they first appear, light or dark orange-color or grayish brown and usually glabrous during their first winter, ultimately becoming ashy gray or light brown. Winter-buds ovoid, obtuse, about ⅛′ long, with light chestnut-brown scales covered, especially near their margins, with loose pale tomentum. Bark ¾′—1′ thick, light gray tinged with red and broken into thick plates separating on the surface into thin irregular appressed scales. Wood heavy, hard, strong, tough, very durable in contact with the ground, rich dark brown, with thick lighter colored sapwood; confounded commercially with the wood of Quercus alba, and used for the same purpose.

Distribution. River swamps and small deep depressions on rich bottom-lands, usually wet throughout the year; southern New Jersey (Riddleton, Salem County), and valley of the Patuxent River, Maryland, southward near the coast to western Florida, through the Gulf states to the valley of the Navasota River, Brazos County, Texas, and through Arkansas to the valley of the Meramec River (Allenton, St. Louis County), Missouri, and to central Tennessee and Kentucky, southern Illinois, and southwestern Indiana to Spencer County; comparatively rare in the Atlantic and east Gulf states; most common and of its largest size in the valley of the Red River, Louisiana, and the adjacent parts of Texas and Arkansas.

Occasionally cultivated in the northeastern states and hardy in eastern Massachusetts.

× Quercus Comptonae Sarg., a hybrid of Quercus lyrata and Q. virginiana, with characters intermediate between those of its parents, discovered many years ago on the banks of Peyton’s Creek, Matagorda County, Texas (now gone), occurs with several individuals near dwellings in Natchez, Adams County, Mississippi, near Selma, Dallas County, Alabama, and in Audubon Park and streets, New Orleans, Louisiana. A tree, sometimes 100° high and one of the handsomest of North American Oaks; also produced artificially by Professor H. Ness by crossing Quercus lyrata and Q. virginiana.

44. Quercus stellata Wang. Post Oak.

Quercus minor Sarg.

Leaves oblong-obovate, usually deeply 5-lobed, with broad sinuses oblique in the bottom, and short wide lobes, broad and truncate or obtusely pointed at apex, gradually narrowed and cuneate, or occasionally abruptly narrowed and cuneate or rounded at base, when they unfold dark red above and densely pubescent, at maturity thick and firm, deep dark green and roughened by scattered fascicled pale hairs above, covered below with gray, light yellow, or rarely silvery white pubescence, usually 4′—5′ long and 3′—4′ across the lateral lobes, with a broad light-colored midrib pubescent on the upper side and tomentose or pubescent on the lower, stout lateral veins arcuate and united near the margins and connected by conspicuous coarsely reticulated veinlets; turning dull yellow or brown in the autumn; petioles stout, pubescent, ½′ to nearly 1′ in length. Flowers: staminate in aments 3′—4′ long; calyx hirsute, yellow, usually divided into 5 ovate acute laciniately cut segments; anthers covered by short scattered pale hairs; pistillate sessile or stalked, their involucral scales broadly ovate, hirsute; stigmas bright red. Fruit sessile or short-stalked; nut oval to ovoid or ovoid-oblong, broad at base, obtuse and naked or covered with pale persistent pubescence at apex, ½′—1′ long, ¼′—¾′ thick, sometimes striate with dark longitudinal stripes, inclosed for one third to one half its length in the cup-shaped, turbinate, or rarely saucer-shaped cup pale and pubescent on the inner surface, hoary-tomentose on the outer surface, and covered by thin ovate scales rounded and acute at apex, reddish brown, and sometimes toward the rim of the cup ciliate on the margins with long pale hairs.

A tree, rarely 100° high, with a trunk 2°—3° in diameter, and stout spreading branches forming a broad dense round-topped head, and stout branchlets coated at first, like the young leaves and petioles, the stalks of the aments of staminate flowers and the peduncles of the pistillate flowers, with thick orange-brown tomentum, light orange color to reddish brown, and covered by short soft pubescence during their first winter, ultimately gray, dark brown, nearly black or bright brown tinged with orange color; usually not more than 50°—60° tall, with a trunk 1°—2° in diameter, and at the northeastern limits of its range generally reduced to a shrub. Winter-buds broadly ovoid, obtuse or rarely acute, ⅛′—¼′ long, with bright chestnut-brown pubescent scales coated toward the margins with scattered pale hairs. Bark ½′—1′ thick, red more or less deeply tinged with brown, and divided by deep fissures into broad ridges covered on the surface with narrow closely appressed or rarely loose scales. Wood very heavy, hard, close-grained, durable in contact with the soil, difficult to season, light or dark brown, with thick lighter colored sapwood; largely used for fuel, fencing, railway-ties, and sometimes in the manufacture of carriages, for cooperage, and in construction.

Distribution. Dry gravelly or sandy uplands; Cape Cod and islands of southern Massachusetts, Rhode Island, Long Island, New York, to western Florida and southern Alabama and Mississippi, and from New York westward to southern Iowa, Missouri, eastern Kansas, western (Dewey County) Oklahoma, Louisiana and Texas; most abundant and of its largest size in the Mississippi basin; ascending on the southern Appalachian Mountains to altitudes of 2500°; the common Oak of central Texas on limestone hills and sandy plains forming the Texas “Cross Timbers”; usually shrubby and rare and local in southern Massachusetts; more abundant southward from the coast of the south Atlantic and the eastern Gulf states to the lower slopes of the Appalachian Mountains; in western Louisiana rarely in the moist soil of low lands.

Showing little variation in the shape of the fruit and in the character of the cup scales Quercus stellata is one of the most variable of North American Oaks in habit, in the nature of the bark, and in the presence or absence of pubescence. Some of the best marked varieties are var. araniosa Sarg., a large tree differing from the type in the usually smooth upper surface of the leaves, in the floccose persistent tomentum on their lower surface, in the less stout usually glabrous yellow or reddish branchlets, and in its scaly bark; dry sandy soil, southern Alabama, western Louisiana, southern Arkansas, eastern Oklahoma and eastern Texas. Var. paludosa Sarg., a tree up to 75° in height, differing from the type in its oblong-obovate leaves 3-lobed above the middle, slightly pubescent branchlets becoming nearly glabrous, and in its scaly bark; in rich deep soil on the often inundated bottoms of Kenison Bayou, near Washington, St. Landry Parish, Louisiana. Var. attenuata Sarg., a large tree differing from the type in the oblong to oblong-obovate narrow leaves 3-lobed at apex and gradually narrowed to the long cuneate base; near Arkansas Post on the White River, Arkansas County, Arkansas. Var. parviloba Sarg., a round-topped tree 25°—30° high, differing from the type in the smaller lobes of the leaves with more prominent reticulate veinlets; dry sandstone hills near Brownwood, Brown County, Texas. Var. anomala Sarg., a tree 15°—18° high, differing from the type in its broadly obovate subcoriaceous leaves slightly 3-lobed and rounded at apex; dry sandstone hills near Brownwood, Brown County, Texas; possibly a hybrid. Var. Palmeri Sarg., a shrub 6°—15° high, forming clumps, differing from the type in its narrow oblong or slightly obovate 5—7-lobed leaves with narrow lobes, densely tomentose below, and in the thicker and more tomentose scales of the cup; sandy uplands, Elk City, Beckham County, Oklahoma. Var. rufescens Sarg., a shrub 12°—15° high, forming large clumps, differing from the type in the rusty brown pubescence on the lower surface of the polymorphous leaves, in the deeper cups of the fruit with thicker basal scales; sandy uplands, Big Spring, Howard County, Texas, and Elk City, Beckham County, Oklahoma. Var. Boyntonii Sarg., a shrub or small tree spreading into thickets, rarely more than 15° in height, differing from the type in its obovate leaves, mostly 3—5-lobed toward the apex, with small rounded lobes, and in their yellow-brown pubescence also found on the branchlets; in glades on the summit of Lookout Mountain, above Gadsden and Attala, Etowah County, Alabama.

The common and most widely distributed of the varieties of the Post Oak is

Quercus stellata var. Margaretta Sarg.

Quercus Margaretta Ashe

Leaves oblong-obovate, rounded at apex, cuneate or rounded at base, 3—5-lobed with usually narrow rounded, but often broad and truncate lobes, the two forms frequently occurring on the same branch, usually becoming glabrous on the upper surface early in the season, slightly pubescent, sometimes becoming nearly glabrous below, 2½′—5′ long and 2′—2½′ wide; petioles glabrous or pubescent. Flowers and Fruit as in the species.

A small tree, rarely 40° high, with slender glabrous reddish or reddish brown branchlets. Winter-buds ovoid, acute, ¼′ long with closely imbricated chestnut-brown scales glabrous, or ciliate on the margins. Bark thick, rough and furrowed, light gray.

Distribution. Usually on dry sandy slopes, hills and ridges, and southward on Pine-barren lands; coast of Virginia (Capron, Southampton County) southward in the coast and middle districts to central (Lake and Orange Counties) and western Florida, through central and southern Alabama, and eastern and southern Mississippi; in Western Louisiana (Natchitoches and Caddo Parishes); southern Arkansas (McNab, Hempstead County), and southwestern Missouri (Prosperity, Jasper County). The common Post Oak of the south Atlantic and Gulf states; occasionally a shrub (f. stonolifera Sarg.) 4°—6° high, with smaller leaves, spreading into broad thickets by stoloniferous shoots; common near Selma, Dallas County, Alabama, and on the dry sand hills of central Oklahoma.

× Quercus Harbisonii Sarg., believed to be a hybrid of Quercus stellata var. Margaretta and Q. virginiana var. geminata, has been found in the neighborhood of Jacksonville, Duval County, Florida.

45. Quercus Garryana Hook. White Oak.

Leaves obovate to oblong, pointed at apex, cuneate or rounded at base, coarsely pinnatifid-lobed, with slightly thickened revolute margins, coated at first with soft pale lustrous pubescence, at maturity thick and firm or subcoriaceous, dark green, lustrous and glabrous above, light green or orange-brown and pubescent or glabrate below, 4′—6′ long, 2′—5′ wide, with a stout yellow midrib, and conspicuous primary veins spreading at right angles, or gradually diverging from the midrib and running to the points of the lobes; sometimes turning bright scarlet in the autumn; petioles stout, pubescent, ½′—1′ in length. Flowers: staminate in hirsute aments; calyx glabrous, laciniately cut into ovate acute slightly ciliate or linear-lanceolate much elongated segments; pistillate sessile and coated with pale tomentum. Fruit sessile or short-stalked; nut oval to slightly obovoid and obtuse, 1′—1¼′ long and ½′—1′ thick, inclosed at the base in a shallow cup-shaped or slightly turbinate cup puberulous and light brown on the inner surface, pubescent or tomentose on the outer, and covered by ovate acute scales with pointed and often elongated tips, thin, free, or sometimes thickened and more or less united toward the base of the cup, decreasing from below upward.

A tree, usually 60°—70° or sometimes nearly 100° high, with a trunk 2°—3° in diameter, stout ascending or spreading branches forming a broad compact head, and stout branchlets coated at first with thick pale rufous pubescence, pubescent or tomentose and light or dark orange color during their first winter, becoming glabrous and rather bright reddish brown in their second year and ultimately gray; frequently at high altitudes, or when exposed to the winds from the ocean, reduced to a low shrub. Winter-buds ovoid, acute, ⅓′—½′ long, densely clothed with light ferrugineous tomentum. Bark ⅛′—1′ thick, divided by shallow fissures into broad ridges separating on the surface into light brown or gray scales sometimes slightly tinged with orange color. Wood strong, hard, close-grained, frequently exceedingly tough, light brown or yellow, with thin nearly white sapwood; in Oregon and Washington used in the manufacture of carriages and wagons, in cabinet-making, shipbuilding, and cooperage, and largely as fuel.

Distribution. Valleys and the dry gravelly slopes of low hills; Vancouver Island and the valley of the lower Fraser River southward through western Washington and Oregon and the California coast-valleys to Marin County; rare and local and the only Oak-tree in British Columbia; abundant and of its largest size in the valleys of western Washington and Oregon; on the islands in the northern part of Puget Sound reduced to a low shrub (Vine Oak); ascending in its shrubby forms to considerable altitudes on the western slopes of the Cascade Mountains; abundant in northwestern California; less common and of smaller size southward.

46. Quercus utahensis Rydb.

Leaves oblong-obovate, gradually narrowed and rounded or cuneate at base, divided often nearly to the midrib by broad or narrow sinuses into four or five pairs of lateral lobes rounded or acute at apex, the upper lobes usually again lobed or undulate, the terminal lobe rounded at apex, entire or three-lobed, thick, dark green, glabrous or nearly glabrous above, pale and soft pubescent below, 2½′—7′ long, 1½′—3½′ wide, with a prominent midrib and primary veins, and conspicuous veinlets; petioles stout, hoary-tomentose early in the season, pubescent or glabrous before maturity, ⅖′—1′ in length. Flowers: staminate in aments covered with fascicled hairs, 2′—2½′ long; calyx scarious, divided to the middle by wide sinuses into narrow acuminate lobes; anthers yellow; pistillate usually solitary or in pairs, the scales of the involucre thickly coated with hoary tomentum. Fruit usually solitary, sessile or raised on a stout pubescent peduncle ¼′—½′ in length; nut ovoid, broad and rounded at the ends, ⅗′—¾′ long, ½′—2½′ thick, usually inclosed for about half its length in the thick hemispheric cup covered with broad ovate pale pubescent scales much thickened on the back and closely appressed below the middle of the cup, gradually reduced in size upward, thin and less closely appressed toward its rim bordered by the free projecting tips of the upper row of scales.

A tree, occasionally 30° high, with a trunk 4′—8′ in diameter, thick erect branches forming a narrow open head, and stout branchlets red-brown and covered with fascicled hairs when they first appear, becoming light orange-brown and puberulous. Bark dark gray-brown, rough and scaly.

Distribution. Dry foothill slopes and the sides of cañons; borders of southwestern Wyoming to the eastern base of the Rocky Mountains of Colorado, and to Utah, northern New Mexico and Arizona, passing into var. mollis Sarg. with thinner scales on the lower part of the cup of the fruit; with the species over its whole range, but most abundant on the Colorado Plateau of northern Arizona; here rarely 40° high, with a trunk 18′—20′ in diameter.

47. Quercus lobata Née. White Oak. Valley Oak.

Leaves oblong to obovate, deeply 7—11 obliquely lobed, rounded at the narrow apex, narrow and cuneate or broad and rounded or cordate at base, the lateral lobes obovate, obtuse or retuse, or ovate and rounded, thin, 2½′—3′ or rarely 4′ long, 1′—2′ wide, dark green and pubescent above, pale and pubescent below, with a stout pale midrib, and conspicuous yellow veins running to the slightly thickened and revolute margins; petioles stout, hirsute, ¼′—½′ in length. Flowers: staminate in hirsute aments 2′—3′ long; calyx light yellow and divided into 6 or 8 acute pubescent ciliate lobes; pistillate solitary, sessile or rarely in elongated few-flowered spikes, their involucral scales broadly ovate, acute, coated with dense pale tomentum, about as long as the narrow calyx-lobes. Fruit solitary or in pairs, nearly sessile; nut conic, elongated, rounded or pointed at apex, 1¼′—2¼′ long, bright green and lustrous when fully grown, becoming bright chestnut-brown, usually inclosed for about one third its length in the cup-shaped cup coated with pale tomentum on the outer surface, usually irregularly tuberculate below, all but the much-thickened basal scales elongated into acute ciliate chestnut-brown free tips longest on the upper scales and forming a short fringe-like border to the rim of the cup.

A tree, often 100° feet high, with a trunk generally 3°—4°, but sometimes 10° in diameter, divided near the ground or usually 20°—30° above it into great limbs spreading at wide angles and forming a broad head of slender branches hanging gracefully in long sprays and sometimes sweeping the ground; less frequently with upper limbs growing almost at right angles with the trunk and forming a narrow rigid head of variously contorted erect or pendant branches, and slender branchlets coated at first with short silky canescent pubescence, ashy gray, light reddish brown, or pale orange-brown and slightly pubescent in their first winter, becoming glabrous and lighter colored during their second year. Winter-buds ovoid, acute, usually about ¼′ long, with orange-brown pubescent scales scarious and frequently ciliate on the margins. Bark ¾′—1½′ thick and covered by small loosely appressed light gray scales slightly tinged with orange or brown, becoming at the base of old trees frequently 5′—6′ thick and divided by longitudinal fissures into broad flat ridges broken horizontally into short plates. Wood hard, fine-grained, brittle, light brown, with thin lighter colored sapwood; used only for fuel.

Distribution. Valleys of western California between the Sierra Nevada and the ocean from the valley of the Trinity River to Kern and Los Angeles (rare) Counties; most abundant and forming open groves in the central valleys of the state.

48. Quercus leptophylla Rydb.

Leaves oblong to oblong-obovate, cuneate or rarely rounded at base, divided about halfway to the midrib into two to four acute or rounded lateral lobes entire or occasionally furnished on the lower side with a small nearly triangular lobe, the terminal lobe short, entire, rounded at apex or three-lobed, when they unfold thickly coated with hoary tomentum, about one-third grown when the flowers open and then covered above with fascicled hairs and tomentose below, at maturity thin, dark green, lustrous and glabrous or nearly glabrous on the upper surface, yellow-green and covered below by short white hairs most abundant on the midrib and veins, 3½′—4′ long, 1½′—2′ wide; petioles slender, pubescent ⅓′—½′ in length. Flowers: staminate in slender villose aments; calyx scarious, divided into five or six narrow acute lobes; anthers dark red-brown as the flowers open; pistillate not seen. Fruit solitary or racemose, sessile or raised on a stout tomentose peduncle ⅖′—⅗′ in length; nut oblong-ovoid, abruptly narrowed and rounded at base, gradually narrowed and rounded at apex, ½′—¾′ long; inclosed for half its length in the thin, hemispheric cup, ⅖′—½′ in diameter, and covered with acuminate only slightly thickened appressed scales densely covered with hoary tomentum.

A tree, 30°—45° high, with a trunk 16′—24′ in diameter, heavy spreading ashy gray branches forming a round-topped head, and stout branchlets, light red-brown or purple and covered with long fascicled hairs when they first appear, becoming light brown and glabrous before autumn. Bark thick, deeply furrowed, covered with small appressed pale gray scales.

Distribution. Rich bottom-lands of the Cucharas River above La Veta, Huerfano County, Colorado; on the Mogollon Mountains, Socorro County, New Mexico.

49. Quercus austrina Small.

Leaves oblong-obovate, acute or rounded at apex, gradually narrowed to the long cuneate base or rarely rounded at base, usually 5-lobed with rounded lobes, the terminal lobe often 3-lobed, the upper lateral lobes pointing forward and much larger than those of the lower pair, or occasionally 3-lobed at the broad apex, or rarely nearly entire with undulate margins, when they unfold sparsely covered below with caducous fascicled hairs, at maturity glabrous, dark green and lustrous above, paler below, 3′—8′ long, 1′—4′ wide, with a prominent midrib and slender primary veins; petioles slender, at first pubescent, soon glabrous, ¼′—⅓′ in length. Flowers not seen. Fruit solitary or in pairs, sessile or raised on a stout stalk up to ½′ in length; nut ovoid, slightly narrowed toward the base, narrowed at the rounded pubescent apex, ½′—¾′ long, ½′ thick, inclosed for a third to a half its length in the thin hemispheric or deep cup-shaped cup, pale tomentose on the inner surface and covered with thin narrow loosely appressed blunt-pointed tomentose scales.

A tree, 70°—80° and rarely 100° high, with a tall trunk 2°—3° in diameter, spreading and ascending branches forming a broad rather open head, and slender glabrous red-brown or gray-brown brittle-jointed branchlets. Winter-buds ovoid to ellipsoid, acute, ⅛′—¼′ long, with closely imbricated acute puberulous chestnut-brown scales ciliate on the margins. Bark pale, scaly, and on old trunks divided into broad ridges.

Distribution. Banks of streams and river bluffs in deep rich soil; coast of South Carolina (Bluffton, Clay County, and near Charleston); Dover, Scriven County, McIntosh County, De Soto Co., Sumter County, and near Bainbridge, Decatur County, Georgia, to central and western Florida (Gainesville, Alachua County, near Santos, Marion County, Lake City, Columbia County, River Junction, Gadsden County, Marianna, Jackson County); western Alabama (Gallion, Hale County, and the neighborhood of Selma [common] and Pleasant Hill, Dallas County); and southern Mississippi (Meridian, Lauderdale County, Laurel, Jones County, Byram and near Jackson, Hinds County, near Natchez, Adams County).

50. Quercus alba L. White Oak.

Leaves oblong-obovate, gradually narrowed and cuneate at base, divided often nearly to the midrib by narrow or broad sinuses usually oblique in the bottom into 7 or 9 lobes, the lateral, narrow, lanceolate or obovate, pointing forward, rounded or acute and often lobed at apex, the terminal usually obovate and 3-lobed, when they unfold bright red above, pale below and coated with soft pubescence, soon becoming silvery white and very lustrous, at maturity thin, firm, glabrous, bright green and lustrous or dull above, pale or glaucous below, 5′—9′ long, 2′—4′ wide, with a stout bright yellow midrib and conspicuous primary veins; turning late in the autumn deep rich vinous red, gradually withering and sometimes remaining on the branches nearly through the winter; petioles stout, glabrous, ½′—1′ in length. Flowers: staminate in hirsute or nearly glabrous aments 2½′—3′ long; calyx bright yellow and pubescent, with acute lobes; pistillate bright red, their involucral scales broadly ovate, hirsute, about as long as the ovate acute calyx-lobes. Fruit sessile or raised on a slender peduncle 1′—2′ long, the two forms sometimes appearing on the same branch; nut ovoid to oblong, rounded at apex, lustrous, ¾′ long, green when fully grown, becoming light chestnut-brown, inclosed for about one fourth its length in the cup-shaped cup coated with pale or light brown tomentum, its scales at the base much thickened, united and produced into short obtuse membranaceous tips, and thinner toward the rim of the cup.

A tree, 80°—100° high, with a trunk 3°—4° in diameter, tall and naked in the forest, short in the open, and surmounted by a broad round-topped head of stout limbs spreading irregularly, small rigid branches, and slender branchlets at first bright green, often tinged with red, and coated with a loose mass of long pale or ferrugineous deciduous hairs, reddish brown during the summer, bright red and lustrous or covered with a glaucous bloom during their first winter, becoming ultimately ashy gray. Winter-buds broadly ovoid, rather obtuse, dark red-brown, about ⅛′ long. Bark light gray slightly tinged with red or brown, or occasionally nearly white, broken into thin appressed scales, becoming on old trunks sometimes 2′ thick and divided into broad flat ridges. Wood strong, very heavy, hard, tough, close-grained, durable, light brown, with thin light brown sapwood; used in shipbuilding, for construction and in cooperage, the manufacture of carriages, agricultural implements, baskets, the interior finish of houses, cabinet-making, for railway-ties and fences, and largely as fuel.

Distribution. Sandy plains and gravelly ridges, rich uplands, intervales, and moist bottom-lands, sometimes forming nearly pure forests; southern Maine to southwestern Quebec, westward through southern Ontario, the southern peninsula of Michigan, southeastern Minnesota, eastern Iowa, and southeastern Nebraska, and southward to western Florida, through the Gulf states to the valley of the Brazos River, Texas and through Arkansas to eastern Oklahoma, eastern Kansas, Missouri, Tennessee, and Kentucky; ascending the southern Appalachian Mountains as a low bush to altitudes of 4500°; most abundant and of its largest size on the lower western slopes of the Alleghany Mountains and on the bottom-lands of the lower Ohio Basin. Passing into

Quercus alba var. latiloba Sarg.

Leaves obovate-oblong, acute or rounded at apex, gradually narrowed and cuneate at base, divided usually less than half way to the midrib into broad rounded lobes; rarely obovate, with undulate margins, or slightly lobed, with broad rounded lobes (var. repanda Michx.). Flowers as in the type. Fruit rarely more than 1½′ in length, with usually thinner cup scales.

Distribution. More abundant than the species and the common northern White Oak.

× Quercus Beadlei Trel., believed to be a hybrid of Quercus alba and Q. Prinus, has been found in a swamp near Clarkton, Bladen County, North Carolina.

× Quercus Bebbiana Schn., probably a hybrid of Quercus alba and Q. macrocarpa, occurs at Charlotte, Chittenden County, Vermont, and near Kenton, Hardin County, Ohio.

× Quercus Deamii Trel., with characters intermediate between those of Quercus alba and Q. Muehlenbergii and evidently a hybrid of these species, is growing near Bluffton, Wells County, Indiana.

× Quercus Faxonii Trel., with characters intermediate between those of Quercus alba and Q. prinoides and evidently a hybrid of these species, has been found in East Walpole, Norfolk County, and Concord, Middlesex County, Massachusetts, and at Greenville, Montcalm County, Michigan.

× Quercus Fernowii Trel., evidently a hybrid of Quercus alba and Q. stellata, has been found near Allenton, St. Louis County, Missouri, and on Red Clay Creek, Virginia.

× Quercus Jackiana Schn., evidently a hybrid of Quercus alba and Q. bicolor, is growing in Franklin Park, Boston.

× Quercus Saulei Schn., with characters intermediate between those of Q. alba and Q. montana and evidently a hybrid of these species, occurs with widely distributed individuals in Vermont (Monkton, Addison County), eastern Massachusetts, near Providence, Rhode Island, New Jersey, eastern Pennsylvania, and the District of Columbia, on the Appalachian Mountains near Biltmore, Buncombe County, and Highlands, Macon County, North Carolina, at Valleyhead, Gadsden County, Alabama, and in Richland County, Illinois.

51. Quercus bicolor Willd. Swamp White Oak.

Quercus platanoides Sudw.

Leaves obovate to oblong-obovate, rounded at the narrowed apex, acute or rounded at the gradually narrowed and cuneate entire base, coarsely sinuate-dentate, or sometimes pinnatifid, with oblique rounded or acute entire lobes, when they unfold light bronze-green and pilose above, covered below with silvery white tomentum, with conspicuous glands on the teeth, at maturity thick and firm, dark green and lustrous on the upper surface, pale or often silvery white or tawny on the lower surface, 5′—6′ long, 2′—4′ wide, with a slender yellow midrib, primary veins running to the points of the lobes, and conspicuous reticulate veinlets; turning in the autumn dull yellow-brown or occasionally orange-color or rarely scarlet before falling; petioles stout, pilose at first, becoming glabrous, ½′—¾′ in length. Flowers: staminate in hairy aments 3′—4′ long; calyx light yellow-green, hirsute with pale hairs, and deeply divided into 5—9 lanceolate acute segments rather shorter than the stamens; pistillate in few-flowered spikes on elongated peduncles covered like the involucral scales with thick white or tawny tomentum; stigmas bright red. Fruit usually in pairs on slender dark brown glabrous puberulous or pubescent stalks 1½′—4′ in length; nut ovoid, with a broad base, rounded, acute and pubescent at apex, light chestnut-brown, ¾′—1¼′ long, ½′—¾′ thick, inclosed for about one third its length in the thick cup-shaped light brown cup pubescent on the inner surface, hoary-tomentose, and sometimes tuberculate or roughened toward the base on the outer surface by the thickened contorted tips of the ovate acute scales, thin, free, acute and chestnut-brown higher on the cup, and often forming a short fringe-like border on its margin, or sometimes entirely covered by thin scales with free acute tips.

A tree, usually 60°—70° or exceptionally 100° high, with a trunk 2°—3° or occasionally 8°—9° in diameter, rather small branches generally pendulous below and rising above into a narrow round-topped open head and often furnished with short pendulous laterals, and stout branchlets, green, lustrous, and slightly scurfy-pubescent when they first appear, light orange color or reddish brown and glabrous or puberulous during their first winter, becoming darker and often purplish and clothed with a glaucous bloom. Winter-buds broadly ovoid and obtuse, or subglobose to ovoid and acute, ⅛′ long, with light chestnut-brown scales usually pilose above the middle. Bark of young stems and small branches smooth, reddish or purplish brown, separating freely into large papery persistent scales curling back and displaying the bright green inner bark; becoming on old trunks 1′—2′ thick, and deeply and irregularly divided by continuous or interrupted fissures into broad flat ridges covered by small appressed gray-brown scales often slightly tinged with red. Wood heavy, hard, strong, tough, light brown, with thin hardly distinguishable sapwood; used in construction, the interior finish of houses, cabinet-making, carriage and boat-building, cooperage, and railway-ties, and for fencing and fuel.

Distribution. Borders of streams and swamps in moist fertile soil; southern Maine to northern Vermont and southwestern Quebec, through Ontario and the southern peninsula of Michigan to southeastern Minnesota, eastern and southern Iowa, southeastern Nebraska and western Missouri, and to the District of Columbia, northern Kentucky and northeastern Oklahoma, and along the Appalachian Mountains to West Virginia; widely scattered, usually in small groves but nowhere very abundant; most common and of its largest size in western New York and northern Ohio.

× Quercus Schuettii Trel., with characters intermediate between those of Quercus bicolor and Q. macrocarpa, and probably a hybrid of these species, occurs at Fort Howard, Brown County, Wisconsin, near Rockfield and Chateaugay, Quebec, and near Rochester and Golah, Munroe County, New York.

52. Quercus Prinus L. Basket Oak. Cow Oak.

Quercus Michauxii Nutt.

Leaves broadly obovate to oblong-obovate, acute or acuminate at apex with a short broad point, cuneate or rounded at the broad or narrow entire base, regularly crenately lobed with oblique rounded entire lobes sometimes furnished with glandular tips, or rarely entire with undulate margins, when they unfold bright yellow-green, lustrous and pubescent above, coated below with thick silvery white or ferrugineous tomentum, at maturity thick and firm or sometimes membranaceous, especially on young and vigorous branches, dark green, lustrous, glabrous or occasionally roughened by scattered fascicled hairs on the upper surface, more or less densely pubescent on the pale green or silvery white lower surface, 6′—8′ long, 3′—5′ wide; turning in the autumn dark rich crimson; petioles stout, ½′—1½′ in length. Flowers: staminate in slender hairy aments, 3′—4′ long; calyx light yellow-green, pilose with long pale hairs, and divided into 4—7 acute lobes; pistillate in few-flowered spikes on short peduncles coated like the involucral scales with dense pale rufous tomentum; stigmas dark red. Fruit solitary or in pairs, sessile or subsessile, or borne on short stout puberulous stalks rarely ½′ in length; nut ovoid to ellipsoidal, with a broad base, and acute, rounded, or occasionally truncate at apex surrounded by a narrow ring of rusty pubescence, or sometimes pilose nearly to the middle, bright brown, rather lustrous, 1′—1½′ long, ¾′—1¼′ thick, inclosed for about one third its length in the thick cup-shaped cup often broad and flat on the bottom, reddish brown and pubescent within, hoary-tomentose and covered on the outer surface by regularly imbricated ovate acute scales rounded and much thickened on the back, their short tips sometimes forming a rigid fringe-like border to the rim of the cup; seed sweet and edible.

A tree, often 100° high, with a trunk sometimes free of branches for 40°—50°, and 3°—7° in diameter, stout branches ascending at narrow angles and forming a round-topped rather compact head, and stout branchlets at first dark green and covered by pale caducous hairs, becoming bright red-brown or light orange-brown during their first winter and ultimately ashy gray. Winter-buds broadly ovoid or oval, acute, ¼′ long, with thin closely and regularly imbricated dark red puberulous scales with pale margins, those of the inner ranks coated on the outer surface with loose pale tomentum. Bark ½′—1′ thick, separating into thin closely appressed silvery white or ashy gray scales more or less deeply tinged with red. Wood heavy, hard, very strong, tough, close-grained, durable, easy to split, light-brown, with thin darker colored sapwood; largely used in all kinds of construction, for agricultural implements, wheels, in cooperage, for fences and fuel, and in baskets.

Distribution. Borders of streams, swamps, and bottom-lands often covered with water; New Jersey (Morristown, Morris County and Pittsgrove, Salem County), near Wilmington, Delaware, southward through the coast and middle districts to Putnam (San Mateo) and Citrus Counties, Florida, through the Gulf states to the valley of the Trinity River, Texas, and through Arkansas and southeastern Missouri to central Tennessee and Kentucky, the valley of the lower Wabash River, Illinois, and southern Indiana eastward to Jefferson County (C. C. Deam); conspicuous from the silvery white bark, the massive trunk, and the broad crown of large bright-colored foliage.

53. Quercus montana L. Chestnut Oak. Rock Chestnut Oak.

Quercus Prinus Engelm., not L.

Leaves obovate or oblong to lanceolate, acute or acuminate or rounded at apex, gradually or abruptly cuneate or rounded or subcordate at the narrow entire base, irregularly and coarsely crenulate-toothed with rounded, acute, or sometimes nearly triangular oblique teeth, when they unfold orange-green or bronze-red, very lustrous, and glabrous above with the exception of the slightly pilose midrib, green and coated below with soft pale pubescence, at maturity thick and firm or subcoriaceous, yellow-green and rather lustrous on the upper surface, paler and covered by fine pubescence on the lower surface, 4½′—9′ long, 1½′—3′ wide, with a stout yellow midrib and conspicuous primary veins, often much broader near the bottom of the tree than on fertile upper branches; turning dull orange color or rusty brown in the autumn; petioles stout or slender, ½′—1′ in length. Flowers: staminate in elongated hirsute aments; calyx light yellow, pilose and deeply divided into 7—9 acute segments tipped with clusters of pale hairs; pistillate in short spikes on stout puberulous dark green peduncles, their involucral scales covered with pale hairs; stigmas dark red. Fruit on short stout stems singly or in pairs; nut ovoid or ellipsoidal, rounded and rather obtuse or pointed at apex, bright chestnut-brown, very lustrous, 1′—1½′ long, ⅝′—1′ thick, inclosed for about half its length or sometimes only at the base in a turbinate or cup-shaped thin cup light brown and pubescent on the inner surface, reddish brown and hoary-pubescent on the outer surface roughened or tuberculate, especially toward the base, by small scales thickened and knob-like with nearly triangular free light brown tips.

A tree, usually 60°—70° or occasionally 100° high, with a trunk 3°—4° or rarely 6°—7° in diameter, divided generally 15° or 20° above the ground into large limbs spreading into a broad open rather irregular head, and stout branchlets green tinged with purple or bronze color and glabrous or pilose when they first appear, light orange color or reddish brown during their first winter, becoming dark gray or brown; on dry exposed mountain slopes often not more than 20°—30° tall, with a trunk 8′—12′ in diameter. Winter-buds ovoid, acute or acuminate, ¼′—½′ long, with bright chestnut-brown scales pilose toward the apex and ciliate on the margins. Bark of young stems and small branches thin, smooth, purplish brown, often lustrous, becoming on old trunks and large limbs ¾′—1½′ thick, dark reddish brown or nearly black, and divided into broad rounded ridges covered with small closely appressed scales. Wood heavy, hard, strong, rather tough, close-grained, durable in contact with the soil, largely used for fencing, railway-ties, and fuel. The bark, which is rich in tannin, is consumed in large quantities in tanning leather.

Distribution. Hillsides and the high rocky banks of streams in rich and deep or sometimes in sterile soil; coast of southern Maine, southern New Hampshire and eastern Massachusetts, southward to Delaware and the District of Columbia, and along the Appalachian Mountains and their foothills to northern Georgia (Wilkes County); ascending to altitudes of 4000°—4500°; in northern Alabama; westward to the shores of Lake Champlain, western New York; southeastern and southern Ohio, and southern Indiana westward to Orange County (C. C. Deam); and to central Kentucky and Tennessee, and northeastern Mississippi (Alcorn, Prentiss and Tishomingo Counties); rare and local in New England and Ontario; abundant on the banks of the lower Hudson River and on the Appalachian hills from southern New York to Alabama; most common and of its largest size on the lower slopes of the mountains of the Carolinas and Tennessee, here often forming a large part of the forest.

× Quercus Sargentii Rehd., believed to be a hybrid of Quercus montana and the European Q. Robur L., has been growing for nearly a hundred years at what is now Holm Lea, Brookline, Norfolk County, Massachusetts.

54. Quercus Muehlenbergii Engelm. Yellow Oak. Chestnut Oak.

Quercus acuminata Sarg.

Leaves usually crowded at the ends of the branches, oblong-lanceolate to broadly obovate, acute or acuminate with a long narrow or with a short broad point, abruptly or gradually narrowed and cuneate or slightly narrowed and rounded or cordate at base, equally serrate with acute and often incurved or broad and rounded teeth tipped with small glandular mucros, or rarely slightly undulate, when they unfold bright bronzy green and puberulous above, tinged with purple and coated below with pale tomentum, at maturity thick and firm, light yellow-green on the upper surface, pale often silvery white and covered with short fine pubescence on the lower surface, 4′—7′ long, 1′—5′ wide, with a stout yellow midrib and conspicuous primary veins running to the points of the teeth; turning in the autumn orange color and scarlet; petioles slender ¾′—1½′ in length. Flowers: staminate in pilose aments 3′—4′ long; calyx light yellow, hairy, deeply divided into 5 or 6 lanceolate ciliate segments; pistillate sessile or in short spikes coated like their involucral scales with thick white tomentum; stigmas bright red. Fruit sessile or raised on a short stout peduncle, solitary or often in pairs; nut broadly ovoid, narrowed and rounded at apex, ½′ to nearly 1′ long, light chestnut-brown, inclosed for about half its length in a thin cup-shaped light brown cup pubescent on the inner, hoary-tomentose on the outer surface, and covered by small obtuse scales more or less thickened and rounded on the back toward the base of the cup, the small free red-brown tips of the upper ranks forming a minute fringe-like border to its rim; seed sweet and sometimes edible.

A tree, 80°—100°, occasionally 160° high, with a tall straight trunk 3°—4° in diameter above the broad and often buttressed base, comparatively small branches forming a narrow shapely round-topped head, and slender branchlets, green more or less tinged with red or purple, pilose when they first appear, light orange color or reddish brown during their first winter, and ultimately gray or brown; east of the Alleghany Mountains and on dry hills often not more than 20°—30° tall. Winter-buds ovoid, acute, ⅛′—¼′ long, with chestnut-brown scales white and scarious on the margins. Bark rarely ½′ thick, broken on the surface into thin loose silvery white scales sometimes slightly tinged with brown. Wood heavy, very hard, strong, close-grained, durable, with thin light-colored sapwood; largely used in cooperage, for wheels, fencing, and railway-ties.

Distribution. Gardner’s Island, Lake Champlain, Vermont, western Massachusetts and Connecticut, near Newberg, Orange County, New York, westward through New York, southern Ontario and southern Michigan to northern Iowa, southeastern Nebraska, eastern Kansas, and Oklahoma to the valley of the Washita River (Garvin County) and to the Devil’s Cañon near Hinton (Caddo County), and southward in the Atlantic states to the District of Columbia, eastern Virginia; sparingly on the eastern foothills of the Blue Ridge in North and South Carolina at altitudes between 1000° and 2000°; in central Tennessee and Kentucky, central and northeastern Georgia, western Florida, and through the Gulf states to the valley of the Guadalupe River, Texas; on the Guadalupe Mountains, Texas, and on the Capitan Mountains, New Mexico (Lincoln County); rare and comparatively local in the Atlantic states, usually on limestone soil; very abundant in the Mississippi basin, growing on ridges, dry flinty hills, deep rich bottom-lands and the rocky banks of streams; probably of its largest size on the lower Wabash River and its tributaries in southern Indiana and Illinois; on the Edwards Plateau (Kemble, Kerr, Uvalde, Bandera and Real Counties), Texas, a form occurs with nuts sometimes 1¼′ long with deeper cups up to 1′ in diameter (var. Brayi Sarg.).

Section 2. Flowers unisexual (usually perfect in Ulmus); calyx regular; stamens as many as its lobes and opposite them; ovary superior, 1-celled (rarely 2-celled in Ulmus); seed 1.

XI. Ulmaceæ.

Trees, with watery juice, scaly buds, terete branchlets prolonged by an upper lateral bud, and alternate simple serrate pinnately veined deciduous stalked 2-ranked leaves unequal and often oblique at base, conduplicate in the bud, their stipules usually fugaceous. Flowers perfect or monœciously polygamous, clustered, or the pistillate sometimes solitary; calyx 4—9-parted or lobed; stamens 4—6; filaments straight; anthers introrse, 2-celled, opening longitudinally; ovary usually 1-celled; ovule solitary, suspended from the apex of the cell, anatropous or amphitropous; styles 2. Fruit a samara, nut, or drupe; albumen little or none; embryo straight or curved; cotyledons usually flat or conduplicate. Five of the thirteen genera of the Elm family occur in North America. Of these four are represented by trees.

Conspectus of the North American Arborescent Genera.

Fruit a dry samara, or nut-like.

Flowers perfect; fruit a samara.

1. Ulmus.

Flowers polygamo-monœcious; fruit nut-like, tuberculate.

2. Planera.

Fruit drupaceous.

Pistillate flowers usually solitary.

3. Celtis.

Pistillate flowers in dichotomous cymes.

4. Trema.

1. Ulmus L. Elm.

Trees, or rarely shrubs, with deeply furrowed bark, branchlets often furnished with corky wings, and buds with numerous ovate rounded chestnut-brown scales closely imbricated in two ranks, increasing in size from without inward, the inner accrescent, replacing the stipules of the first leaves, deciduous, marking the base of the branchlet with persistent ring-like scars. Leaves simply or doubly serrate; stipules linear, lanceolate to obovate, entire, free or connate at base, scarious, inclosing the leaf in the bud, caducous. Flowers from axillary buds near the ends of the branches similar to but larger than the leaf-buds, the outer scales sterile, the inner bearing flowers and rarely leaves. Flowers perfect, jointed on slender bibracteolate pedicels from the axils of linear acute scarious bracts, in pedunculate or subsessile fascicles or cymes sometimes becoming racemose, appearing in early spring before the leaves in the axils of those of the previous year, or autumnal in the axils of leaves of the year; calyx campanulate, 5—9-lobed, membranaceous, marcescent; stamens 5 or 6 inserted under the ovary; filaments filiform or slightly flattened, erect in the bud, becoming exserted; anthers oblong, emarginate, and subcordate; ovary sessile or stipitate, compressed, crowned by a simple deeply 2-lobed style, the spreading lobes papillo-stigmatic on the inner face, usually 1-celled by abortion, rarely 2-celled; ovule amphitropous; micropyle extrorse, superior. Fruit an ovoid or oblong, often oblique, sessile or stipitate samara surrounded at base by the remnants of the calyx, the seminal cavity compressed, slightly thickened on the margin, chartaceous, produced into a thin reticulate-venulose membranaceous light brown broad or rarely narrow wing naked or ciliate on the margin, tipped with the remnants of the persistent style, or more or less deeply notched at apex, and often marked by the thickened line of the union of the two carpels. Seed ovoid, compressed, without albumen, marked on the ventral edge by the thin raphe; testa membranaceous, light or dark chestnut-brown, of two coats, rarely produced into a narrow wing; embryo erect; cotyledons flat or slightly convex, much longer than the superior radicle turned toward the oblong linear pale hilum.

Ulmus, with eighteen or twenty species, is widely distributed through the boreal and temperate regions of the northern hemisphere with the exception of western North America, reaching in the New World the mountains of southern Mexico and in the Old World the Sikkim Himalaya, western China, and Japan. Of the exotic species, Ulmus procera Salisb., the so-called English Elm, and Ulmus glabra, Huds., the Scotch Elm, and several of its varieties, have been largely planted for shade and ornament in the north Atlantic states, where old and large specimens of the former can be seen, especially in the neighborhood of Boston.

Ulmus produces heavy, hard, tough, light-colored wood, often difficult to split. The tough inner bark of some of the species is made into ropes or woven into coarse cloth, and in northern China nourishing mucilaginous food is prepared from the inner bark.

Ulmus is the classical name of the Elm-tree.

Conspectus of the North American Species.

Flowers vernal, appearing before the leaves.

Flowers on slender drooping pedicels; fruit ciliate on the margins.

Wing of the fruit broad.

Bud-scales and fruit glabrous; branchlets destitute of corky wings; leaves obovate-oblong to elliptic, usually smooth on the upper, soft-pubescent on the lower surface.

1. U. americana (A, C).

Bud-scales puberulous; branches often furnished with corky wings; fruit hirsute; leaves obovate to oblong, smooth on the upper, soft-pubescent on the lower surface.

2. U. racemosa (A).

Wing of the fruit narrow; bud-scales glabrous or slightly puberulous; branchlets usually furnished with broad corky wings; fruit hirsute, leaves ovate-oblong to oblong-lanceolate, smooth on the upper, soft-pubescent on the lower surface.

3. U. alata (A, C).

Flowers on short pedicels; fruit naked on the margins; bud-scales coated with rusty hairs; fruit pubescent; leaves ovate-oblong, scabrous on the upper, pubescent on the lower surface.

4. U. fulva (A, C).

Flowers autumnal, appearing in the axils of leaves of the year; branchlets furnished with corky wings; fruit hirsute.

Bud-scales puberulous; flowers on short pedicels; leaves ovate, scabrous on the upper, soft-pubescent on the lower surface.

5. U. crassifolia (C).

Bud-scales glabrous; flowers on long pedicels; leaves oblong to oblong-obovate, acuminate, glabrous on the upper, pale and puberulous on the lower surface.

6. U. serotina (C).

1. Ulmus americana L. White Elm.

Leaves obovate-oblong to elliptic, abruptly narrowed at apex into a long point, full and rounded at base on one side and shorter and cuneate on the other, coarsely doubly serrate with slightly incurved teeth, when they unfold coated below with pale pubescence and pilose above with long scattered white hairs, at maturity 4′—6′ long, 1′—3′ wide, dark green and glabrous or scabrate above, pale and soft-pubescent or sometimes glabrous below, with a narrow pale midrib and numerous slender straight primary veins running to the points of the teeth and connected by fine cross veinlets; turning bright clear yellow in the autumn before falling; petioles stout, ¼′ in length; stipules linear-lanceolate, ½′—2′ long. Flowers on long slender drooping pedicels sometimes 1′ in length, in 3 or 4-flowered short-stalked fascicles; calyx irregularly divided into 7—9 rounded lobes ciliate on the margins, often somewhat oblique, puberulous on the outer surface, green tinged with red above the middle; anthers bright red; ovary light green, ciliate on the margins with long white hairs; styles light green. Fruit on long pedicels in crowded clusters, ripening as the leaves unfold, ovoid to obovoid-oblong, slightly stipitate, conspicuously reticulate-venulose, ½′ long, ciliate on the margins, the sharp points of the wings incurved and inclosing the deep notch.

A tree, sometimes 100°—120° high, with a tall trunk 6°—11° in diameter, frequently enlarged at the base by great buttresses, occasionally rising with a straight undivided shaft to the height of 60°—80° and separating into short spreading branches, more commonly divided 30°—40° from the ground into numerous upright limbs gradually spreading and forming an inversely conic round-topped head of long graceful branches, often 100° or rarely 150° in diameter, and slender branchlets frequently fringing the trunk and its principal divisions, light green and coated at first with soft pale pubescence, becoming in their first winter light reddish brown, glabrous or sometimes puberulous and marked by scattered pale lenticels, and by large elevated semiorbicular leaf-scars showing the ends of three large equidistant fibro-vascular bundles, later becoming dark reddish brown and finally ashy gray. Winter-buds ovoid, acute, slightly flattened, about ⅛′ long, with broadly ovate rounded light chestnut-brown glabrous scales, the inner bright green, ovate, acute, becoming on vigorous shoots often nearly 1′ in length. Bark 1′—1½′ thick, ashy gray, divided by deep fissures into broad ridges separating on the surface into thin appressed scales. Wood heavy, hard, strong, tough, difficult to split, coarse-grained, light brown, with thick somewhat lighter colored sapwood; largely used for the hubs of wheels, saddle-trees, in flooring and cooperage, and in boat and shipbuilding.

Distribution. River-bottom lands, intervales, low rich hills, and the banks of streams; southern Newfoundland to the northern shores of Lake Superior and the headwaters of the Saskatchewan, southward to the neighborhood of Lake Istokpoga, De Soto County, Florida, westward in the United States to the Turtle Mountains of North Dakota, the Black Hills of South Dakota, western Nebraska, central Kansas and Oklahoma, and the valley of the upper Colorado River (Fort Chadbourne, Coke County), Texas; very common northward, less abundant and of smaller size southward; abundant on the banks of streams flowing through the midcontinental plateau.

Largely planted as an ornamental and shade tree in the northern states, and rarely in western and northern Europe.

2. Ulmus racemosa Thomas. Rock Elm. Cork Elm.

Ulmus Thomasii Sarg.

Leaves obovate to oblong-oval, rather abruptly narrowed at apex into a short broad point, equally or somewhat unequally rounded, cuneate or subcordate at base, and coarsely doubly serrate, when they unfold pilose on the upper surface and covered on the lower with soft white hairs, at maturity 2′—2½′ long, ¾′—1′ wide, thick and firm, smooth, dark green and lustrous above, paler and soft-pubescent below, especially on the stout midrib and the numerous straight veins running to the point of the teeth and connected by obscure cross veinlets; turning in the autumn bright clear yellow; petioles pubescent, about ¼′ in length; stipules ovate-lanceolate, conspicuously veined, light green, marked with dark red on the margins above the middle, ⅔′ long, clasping the stem by their abruptly enlarged cordate base conspicuously dentate with 1—3 prominent teeth on each side, falling when the leaves are half grown. Flowers on elongated slender drooping pedicels often ½′ long, in 2—4, usually in 3-flowered, puberulous cymes becoming more or less racemose by the lengthening of the axis of the inflorescence, and when fully grown sometimes 2′ in length; calyx green, divided nearly to the middle into 7 or 8 rounded dark red scarious lobes; anthers dark purple; ovary coated with long pale hairs most abundant on the margins; styles light green. Fruit ripening when the leaves are about half grown, ovoid or obovoid-oblong, ½′ long, with a shallow open notch at the apex, obscurely veined, pale pubescent, ciliate on the slightly thickened border of the broad wing, the margin of the seminal cavity scarcely thickened.

A tree, 80°—100° high, with a trunk occasionally 3° in diameter, and often free of branches for 60°, short stout spreading branches forming a narrow round-topped head, and slender rigid branchlets, light brown when they first appear, and coated with soft pale pubescence often persistent until their second season, becoming light reddish brown, puberulous or glabrous and lustrous in their first winter, and marked by scattered oblong lenticels and large orbicular or semiorbicular leaf-scars displaying an irregular row of 4—6 fibro-vascular bundle-scars, ultimately dark brown or ashy gray, and usually furnished with 3 or 4 thick corky irregular wings often ½′ broad, and beginning to appear in their first or more often during their second year. Winter-buds ovoid, acute, ¼′ long, with broadly ovate rounded chestnut-brown scales pilose on the outer surface, ciliate on the margins, the inner scales becoming ovate-oblong to lanceolate, and ½′ long, often dentate at the base, with 1 or 2 minute teeth on each side, bright green below the middle, marked with a red blotch above, and white and scarious at the apex. Bark ¾′—1′ thick, gray tinged with red, and deeply divided by wide irregular interrupted fissures into broad flat ridges broken on the surface into large irregularly shaped scales. Wood heavy, hard, very strong and tough, close-grained, light clear brown often tinged with red, with thick lighter colored sapwood; largely employed in the manufacture of many agricultural implements, for the framework of chairs, hubs of wheels, railway-ties, the sills of buildings, and other purposes demanding toughness, solidity and flexibility.

Distribution. Dry gravelly uplands, low heavy clay soils, rocky slopes and river cliffs; Province of Quebec westward through Ontario, the southern peninsula of Michigan and central Wisconsin to northeastern Nebraska, western Missouri and eastern Kansas, and southward to northern New Hampshire, southern Vermont, western New York, (valley of the Genesee River), northern New Jersey, southern Ohio (near Columbus, Franklin County), and central Indiana; rare in the east and toward the extreme western and southern limits of its range.

Occasionally planted as a shade and ornamental tree in the northern states.

3. Ulmus alata Michx. Wahoo. Winged Elm.

Leaves ovate-oblong to oblong-lanceolate, often somewhat falcate, acute or acuminate, unequally cuneate or rounded or subcordate at base, and coarsely doubly serrate with incurved teeth, when they unfold pale green often tinged with red, coated on the lower surface with soft white pubescence and glabrous or nearly so on the upper surface, at maturity thick and firm or subcoriaceous, dark green and smooth above, pale and soft-pubescent below, especially on the stout yellow midrib and numerous straight prominent veins often forked near the margins of the leaf and connected by rather conspicuous reticulate veinlets; turning yellow in the autumn; their petioles stout, pubescent, ⅓′ in length; stipules linear-obovate, thin and scarious, tinged with red above the middle, often nearly 1′ long. Flowers on drooping pedicels, in short few-flowered fascicles; calyx glabrous and divided nearly to the middle into 5 broad ovate rounded lobes as long as the hoary-tomentose ovary raised on a short slender stipe. Fruit ripening before or with the unfolding of the leaves, oblong, ⅓′ in length, contracted at base into a long slender stalk, gradually narrowed and tipped at apex with long incurved awns, and covered with long white hairs most numerous on the thickened margin of the narrow wing; seed ovoid, pointed, ⅛′ long, pale, chestnut-brown, slightly thickened into a narrow wing-like margin.

A tree, occasionally 80°—100° but usually not more than 40°—50° high, with a trunk 2°—3° in diameter, short stout straight or erect branches forming a narrow oblong rather open round-topped head, and slender branchlets glabrous or puberulous and light green tinged with red when they first appear, becoming light reddish brown or ashy gray and glabrous, or on vigorous individuals frequently pilose in their first winter, marked by occasional small orange-colored lenticels and by small elevated horizontal semiorbicular leaf-scars, sometimes naked, more often furnished with usually 2 thin corky wings beginning to grow during their first or more often during their second season, abruptly arrested at the nodes, often ½′ wide, and persistent for many years. Winter-buds slender, acute, ⅛′ long, dark chestnut-brown, with glabrous or puberulous scales, those of the inner ranks becoming oblong or obovate, rounded and tipped with a minute mucro, thin and scarious, light red, especially above the middle, and ½′ long. Bark rarely exceeding ¼′ in thickness, light brown tinged with red, and divided by irregular shallow fissures into flat ridges covered by small closely appressed scales. Wood heavy, hard, not strong, close-grained, difficult to split, light brown, with thick lighter colored sapwood; sometimes employed for the hubs of wheels and the handles of tools. Ropes used for fastening the covers of cotton bales are sometimes made from the inner bark.

Distribution. Usually on dry gravelly uplands, less commonly in alluvial soil on the borders of swamps and the banks of streams, and occasionally in inundated swamps; southeastern Virginia, southwestern Indiana, southern Illinois (Richland and Johnson Counties) and southern Missouri, and southward to central Florida (Lake County), and the valley of the Guadalupe River, Texas; ranging westward in Oklahoma to Garfield County (near Kingfisher, G. W. Stevens).

Often planted as a shade-tree in the streets of towns and villages of the southern states.

4. Ulmus fulva Michx. Slippery Elm. Red Elm.

Leaves ovate-oblong, abruptly contracted into a long slender point, rounded at base on one side and short-oblique on the other, and coarsely doubly serrate with incurved callous-tipped teeth, when they unfold thin, coated below with pale pubescence, pilose above with scattered white hairs, at maturity thick and firm, dark green and rugose with crowded sharp-pointed tubercles pointing toward the apex of the leaf above, soft, smooth, and coated below, especially on the thin midrib and in the axils of the slender straight veins with white hairs, 5′—7′ long, 2′—3′ wide; turning a dull yellow color in the autumn; petioles stout, pubescent, ⅓′ in length; stipules obovate-oblong to oblong-lanceolate, thin and scarious, pale-pubescent, and tipped with clusters of rusty brown hairs. Flowers on short pedicels, in crowded fascicles; calyx green, covered with pale hairs, divided into 5—9 short rounded thin equal lobes; stamens with slender light yellow slightly flattened filaments and dark red anthers; stigmas slightly exserted, reddish purple, papillose with soft white hairs. Fruit ripening when the leaves are about half grown, semiorbicular, rounded and bearing the remnants of the styles or slightly emarginate at apex, rounded or cuneate at base, ½′ broad, the seminal cavity coated with thick rusty brown tomentum, the broad thin wing obscurely reticulate-veined, naked on the thickened margin, and marked by the dark conspicuous horizontal line of union of the two carpels; seed ovoid, with a large oblique pale hilum, a light chestnut-brown coat produced into a thin border wider below than above the middle of the seed.

A tree, 60°—70° high, with a trunk occasionally 2° in diameter, spreading branches forming a broad open flat-topped head, and stout branchlets bright green, scabrate, and coated with soft pale pubescence when they first appear, becoming light brown by midsummer, often roughened by small pale lenticels, and in their first winter ashy gray, orange color or light red-brown, and marked by large elevated semiorbicular leaf-scars showing the ends of 3 conspicuous equidistant fibro-vascular bundles, ultimately dark gray or brown. Winter-buds ovoid, obtuse, ¼′ long, with about 12 scales, the outer broadly ovate, rounded, dark chestnut-brown, and covered by long scattered rusty hairs, the inner when fully grown ½′ long, ⅛′—¼′ wide, light green, strap-shaped, rounded and tipped at the apex with tufts of rusty hairs, puberulous on the outer surface, slightly ciliate on the margins, gradually growing narrower and passing into the stipules of the upper leaves. Bark frequently 1′ thick, dark brown tinged with red, divided by shallow fissures and covered by large thick appressed scales. Wood heavy, hard, strong, very close-grained, durable, easy to split, dark brown or red, with thin lighter colored sapwood; largely used for fence-posts, railway-ties, the sills of buildings, the hubs of wheels, and in agricultural implements. The thick fragrant inner bark is mucilaginous and demulcent, and is employed in the treatment of acute febrile and inflammatory affections.

Distribution. Banks of streams and low rocky hillsides in deep rich soil; comparatively common in the valley of the St. Lawrence River, Province of Quebec, and through Ontario to northern and eastern South Dakota, northeastern and eastern Nebraska, southeastern Kansas, and Oklahoma to the valley of the Canadian River (McClain County), and southward to western Florida, central Alabama and Mississippi, western Louisiana and the valley of the upper Guadalupe (Kerr County) and Leon Rivers (Comal County), Texas; in the South Atlantic states not common and mostly confined to the middle districts, ascending to altitudes of 2000° on the southern Appalachian foothills.

5. Ulmus crassifolia Nutt. Cedar Elm.

Leaves elliptic to ovate, acute or rounded at apex, unequally rounded or cuneate and often oblique at base, coarsely and unequally doubly serrate with callous-tipped teeth, when they unfold thin, light green tinged with red, pilose above and covered below with soft pale pubescence, at maturity thick and subcoriaceous, dark green, lustrous and roughened by crowded minute sharp-pointed tubercles on the upper surface and soft pubescent on the lower surface, 1′—2′ long, ½′—1′ wide, with a stout yellow midrib, and prominent straight veins connected by conspicuous more or less reticulate cross veinlets; usually turning bright yellow late in the autumn; petioles stout, tomentose, ¼′—½′ in length; stipules ½′ long, linear-lanceolate, red and scarious above, clasping the stem by their green and hairy bases, deciduous when the leaves are about half grown. Flowers usually opening in August and sometimes also in October, on slender pedicels ⅓′—½′ long and covered with white hairs, in 3—5-flowered pedunculate fascicles; calyx divided to below the middle into oblong pointed lobes hairy at base; ovary hirsute, crowned with two short slightly exserted stigmas. Fruit ripening in September and rarely also in November, oblong, gradually and often irregularly narrowed from the middle to the ends, short-stalked, deeply notched at apex, ⅓′ to nearly ½′ long, covered with soft white hairs, most abundant on the slightly thickened margin of the broad wing; seed oblique, pointed, and covered by a dark chestnut-brown coat.