Division II. Dicotyledons (part 4)

Leaves oblong-obovate, acute or occasionally rounded at apex, gradually narrowed to the long cuneate base, and finely serrate above the middle with straight or incurved glandular teeth, more than half grown and sparingly villose on the upper side of the midrib when the flowers open the middle of April, and at maturity glabrous, dark green and lustrous above, paler below, 1′—1½′ long, and ½′—¾′ wide, with a thin midrib, and slender primary veins mostly within the parenchyma; petioles slender, glabrous, ⅓′—½′ in length; leaves at the end of vigorous shoots broad-obovate, rounded or acute at apex, coarsely serrate, 2′—2½′ long, and 1′—1¼′ wide. Flowers ⅖′—½′ in diameter, on slender pedicels, in mostly 5—8-flowered glabrous corymbs; calyx-tube narrowly obconic, the lobes short and broad, acuminate, entire or slightly dentate near the middle, sparingly villose on the inner surface; stamens 20; anthers small, nearly white; styles 2 or 3. Fruit on slender drooping pedicels, short-oblong, rounded at the ends, dull red, about ½′ long and ⅓′ thick; calyx prominent, with reflexed closely appressed persistent lobes; flesh thin, dry and hard; nutlets 2 or 3, broad and rounded at base, narrowed at apex, about ¼′ long.

A tree, 18°—20° high, with a slender stem covered with close dark slightly ridged bark, small wide-spreading branches forming a flat-topped head, and slender slightly zigzag orange or red-brown branchlets unarmed, or armed with few or many straight or slightly curved dark chestnut-brown shining spines ½′—1′ in length.

Distribution. Woods in low sandy soil; eastern Texas (near Marshall, Harrison County, and Livingston, Polk County).

15. Cratægus Engelmannii Sarg.

Leaves oblong-obovate or rarely elliptic, rounded or often short-pointed and acute at apex, gradually narrowed or entire below, finely crenulate-serrate usually only above the middle and generally only at the apex, nearly fully grown and roughened on the upper surface by short rigid pale hairs when the flowers open about the middle of May, and at maturity coriaceous, dark green, lustrous and scabrate above, pale below, and pilose on both surfaces of the slender midrib and obscure primary veins and veinlets, 1′—1½′ long, and ½′—1′ wide; petioles glandular, villose when they first appear, soon glabrous, usually about ¼′ in length. Flowers ¾′ in diameter, on slender pedicels, in broad loose 8—11-flowered villose corymbs; calyx-tube narrowly obconic, villose or nearly glabrous, the lobes narrow, acuminate, entire, glabrous on the outer surface, usually puberulous on the inner surface; stamens 10; anthers small, rose color; styles 2 or 3. Fruit ripening early in November, on slender pedicels, in drooping many-fruited glabrous clusters, globose or short-oblong, bright orange-red, with a yellow cheek, about ⅓′ in diameter; calyx prominent, with large spreading lobes usually deciduous before the fruit ripens; nutlets 2 or 3, thick, with a broad rounded ridge, ¼′ long.

A tree, 15°—20° high, with a trunk 5′—6′ in diameter, wide-spreading usually horizontal branches forming a low flat-topped or rounded head, and branchlets covered with long pale hairs when they first appear, soon glabrous and bright red-brown, becoming gray or gray tinged with red during their second year, and armed with numerous stout straight or slightly curved spines 1½′—2½′ long.

Distribution. Dry limestone slopes and ridges; common near Allenton and Pacific, St. Louis and Franklin counties, Missouri; near Eureka Springs, Carroll County, Arkansas.

16. Cratægus montivaga Sarg.

Leaves obovate to oval, rhombic or suborbicular, rounded, acute or acuminate or abruptly short-pointed at apex, concave-cuneate at base, and sharply coarsely serrate usually to below the middle with straight acuminate glandular teeth, covered above with short white hairs and glabrous below when they unfold, and at maturity dark green, lustrous and scabrate above, pale yellow-green below, 1′—1¼′ long, and ¾′—1′ wide, with a slender midrib and prominent primary veins; petioles slender, villose early in the season, becoming glabrous, about ¼′ in length. Flowers opening late in April, about ½′ in diameter, on villose pedicels ¼′—½′ long, in compact mostly 7—10-flowered villose corymbs, their bracts and bractlets linear-obovate, conspicuously glandular-serrate; calyx-tube broadly obconic, glabrous or with occasional hairs near the base, the lobes gradually narrowed from a wide base, glandular-serrate, sometimes laciniate near the acuminate apex, glabrous on the outer surface, villose on the inner surface; stamens 10—15, usually 10; anthers pink; styles 2 or 3. Fruit ripening late in September or in October, on erect nearly glabrous or villose pedicels, short-oblong to ellipsoid, orange-red, about ⅓′ long; the calyx enlarged and conspicuous; flesh thin, yellow-green; nutlets 2 or 3, rounded at apex, with a low broad rounded ridge, about ¼′ long.

A bushy tree, rarely more than 12°—15° high, with a short trunk 10′—12′ in diameter, erect and spreading branches, and slender nearly straight branchlets orange-brown and covered with long scattered pale hairs when they first appear, dull red-brown and glabrous at the end of their first season, becoming gray the following year. Bark of the branches smooth and dark brown, becoming slightly scaly on the trunk.

Distribution. Rocky banks of streams; western Texas (Comal, Kendall, Bandera, Edwards, Brown and Calhoun Counties, and on the Davis Mountains, Jeff Davis County); common on the banks of the Guadalupe and other streams on the Edwards Plateau.

Interesting as the extreme southwestern representative of the Crus-galli Group, and its only species in western Texas.

17. Cratægus denaria Beadl.

Leaves oval, oblong-obovate or elliptic, acute or acuminate at apex, gradually narrowed from near the middle and cuneate and entire below, and coarsely often doubly serrate above with straight teeth, when they unfold tinged with red and slightly pilose above and glabrous below, nearly fully grown when the flowers open toward the end of May, and at maturity firm to subcoriaceous, bright green and lustrous on the upper surface, pale on the lower surface, 2½′—3′ long, and ¾′—1¼′ wide, with a slender midrib and few remote thin primary veins; turning in the autumn orange, yellow, or brown; petioles stout, conspicuously glandular, and about ¼′ in length; leaves at the end of vigorous shoots broadly oval to ovate or obovate, occasionally incisely lobed, 2½′—3′ long, and 1½′—2′ wide. Flowers ½′—⅔′ in diameter, on long slender pedicels, in broad lax many-flowered sparingly villose corymbs; calyx narrowly obconic, glabrous, the lobes slender, elongated, acuminate and glandular at apex, mostly entire or slightly serrate below; stamens usually 10; styles 3—5. Fruit on long slender pedicels, in drooping few-fruited clusters, globose to subglobose, ¼′—/16′ in diameter, orange-red, the calyx somewhat enlarged, with spreading or closely appressed lobes; nutlets 3—5, slightly ridged on the back, about /16′ long.

A tree, 18°—20° high, with a trunk sometimes 8′ in diameter, spreading branches, and branchlets sparingly villose with long matted white hairs when they first appear, soon glabrous, and unarmed or armed with occasional straight slender spines about 1½′ long.

Distribution. Banks of streams, eastern Mississippi; common in the neighborhood of Columbus, Lowndes County.

18. Cratægus signata Beadl.

Leaves obovate to elliptic, rounded and often short-pointed or acute at apex, gradually narrowed from near the middle and cuneate at the entire base, and sharply glandular-serrate usually only above the middle, about half grown when the flowers open early in April, and then gray-green and coated above and on the lower side of the midrib and principal veins with short pale hairs, and at maturity thin and firm in texture, dark green, lustrous and slightly pilose above, paler and pubescent below on the slender midrib and 2—5 pairs of primary veins, 1½′—2′ long, and ¾′—1′ wide; petioles slender, grooved above, glandular, usually about ⅓′ in length; leaves at the end of vigorous shoots often broad-ovate to elliptic, coarsely dentate or sometimes incisely lobed, frequently 2½′ long and 2′ wide. Flowers about ¾′ in diameter, on slender pedicels, in few-flowered compact hairy corymbs; calyx-tube narrowly obconic, villose with long matted hairs, the lobes narrow, acute, entire or irregularly glandular-serrate, usually glabrous on the outer surface, villose on the inner surface; stamens 10; styles 3—5, surrounded at base by a few pale hairs. Fruit ripening and falling toward the end of October, in few-fruited drooping slightly villose clusters, short-oblong, rounded at the ends, dark red, more or less pruinose, marked by numerous pale dots, and about ½′ long; calyx enlarged, with elongated closely appressed lobes usually persistent on the ripe fruit; flesh thin and yellow; nutlets 3—5, prominently ridged and grooved on the back, about ¼′ long.

A tree, usually 15°—18° high, with a tall trunk 4′—5′ in diameter, covered with ashy gray bark, often nearly black near the base of old stems, and separating freely into thin plate-like scales, numerous ascending or spreading branches forming a round-topped or oval compact head, and stout chestnut-brown branchlets armed with stout, nearly straight bright chestnut-brown spines 1′—2′ long.

Distribution. Open glades and dry copses of the Pine-covered coast-plain of southern Alabama.

19. Cratægus edita Sarg.

Leaves oblong-obovate or rarely elliptic, acute at the gradually narrowed apex, gradually narrowed from near the middle to the cuneate entire base, and coarsely and often doubly serrate above, when the flowers open from the 15th to the 20th of April lustrous and scabrate on the upper surface with short rigid pale hairs and puberulous on the lower surface, and at maturity coriaceous, dark green, lustrous, and slightly roughened above, pale yellow-green and scabrate below, 1½′—2′ long, and ½′—1′ wide; petioles stout, villose, becoming pubescent or puberulous, ⅓′—½′ in length; leaves at the end of vigorous shoots often slightly divided into lateral lobes, more coarsely serrate and sometimes 3′ long, and 1½′ wide, with stout broadly winged petioles. Flowers ½′—⅔′ in diameter, on slender villose pedicels, in villose few-flowered narrow corymbs; calyx-tube narrowly obconic, glabrous or slightly hairy toward the base, the lobes linear-lanceolate, usually entire or obscurely glandular-serrate, glabrous on the outer surface and puberulous on the inner surface; stamens 20; anthers small, rose color; styles 2 or 3. Fruit ripening early in October or in November, on stout glabrous or slightly villose pedicels usually about ½′ long, in drooping few-fruited clusters, short-oblong, rounded at the ends, slightly pruinose, dull green tinged with red, ¼′—⅓′ long, with a prominent calyx-tube and elongated spreading lobes puberulous on the inner surface and often deciduous before the ripening of the fruit; flesh very thin, green, dry and hard; nutlets 2 or 3, with a broad low rounded ridge, ¼′ long.

A tree, in low moist ground sometimes 40° high, with a trunk 1° in diameter, free of branches for 18°—20°, stout horizontal branches forming a broad round symmetrical head, and nearly straight branchlets villose when they first appear, soon glabrous, light chestnut-brown becoming dark gray-brown in their second or third year, and armed with stout or slender straight chestnut-brown spines 1′—2′ long; or on the dry soil of low hills much smaller and generally 20°—25° high.

Distribution. Low wet woods on the borders of streams, and on dry hills in forests of Oak and Pine; near Marshall, Harris County, Texas; Natchitoches, Natchitoches Parish, Louisiana.

20. Cratægus tersa Beadl.

Leaves oblong to obovate, rounded and obtuse at apex, gradually narrowed to the concave-cuneate entire base, and coarsely serrate above with acute or rounded teeth, when they unfold tinged with red, sparingly villose above and tomentulose below, nearly fully grown when the flowers open the middle of April, and at maturity coriaceous, dark green, lustrous, and glabrous or scabrate above, pale and pubescent below, 1½′—2′ long, and 1′—1¼′ wide, with a slender midrib and thin primary veins; turning in the autumn yellow, orange, and brown; petioles stout, at first hoary-tomentose, glabrous at maturity, about ½′ in length; leaves on the end of vigorous shoots, broad-obovate, short-pointed at the rounded apex, often 2′ long and 1½′ wide, with a prominent midrib and primary veins. Flowers ⅝′—¾′ in diameter, on short stout hairy pedicels, in usually 8—10-flowered very compact corymbs densely clothed with long matted pale hairs; calyx-tube narrowly obconic, villose, the lobes acuminate, glandular-serrate, villose on the outer and slightly pilose on the inner surface; stamens 18—20; anthers pale rose color, styles usually 2 or 3. Fruit ripening in October, on stout glabrous stems, in compact drooping few-fruited clusters, globose to subglobose or short-oblong, about ⅜′ long, dark red; calyx prominent, with enlarged erect or spreading glandular-serrate lobes; flesh thin, yellow, dry and mealy; nutlets 2 or 3, mostly obtuse and rounded at the ends, about ¼′ long.

A tree, sometimes 18°—20° high, with a trunk 6′—8′ in diameter, spreading branches forming a broad flat-topped head, and stout chestnut-brown branchlets at first pilose, becoming glabrous before autumn, and usually unarmed.

Distribution. Low woods west of Opelousas, St. Landry Parish, Louisiana.

21. Cratægus berberifolia T. & G.

Leaves oblong-obovate to elliptic, rounded or gradually narrowed at apex, narrowed from above the middle to the cuneate entire base, and serrate above with straight or incurved teeth, nearly fully grown when the flowers open at the end of March or early in April and then roughened above by short rigid white hairs, and whitish and pubescent below, and at maturity subcoriaceous, dark green, lustrous and nearly glabrous on the upper surface, pale and pubescent on the lower surface especially on the thin midrib and slender primary veins, 1½′—2′ long, and ¾′—1′ wide; petioles comparatively slender, at first densely villose, becoming glabrous, usually about ½′ in length. Flowers ½′—⅔′ in diameter, on slender villose pedicels, in compact mostly 4—5-flowered villose corymbs; calyx-tube narrowly obconic, thickly coated with long matted pale hairs, the lobes slender, acuminate, sparingly villose or nearly glabrous on the outer surface, villose on the inner surface, entire or slightly serrate; stamens 20; anthers yellow; styles 2 or 3, surrounded at base by a narrow ring of pale hairs. Fruit ripening early in October, on slender pedicels, in few-fruited drooping puberulous clusters, subglobose, orange with a red cheek, about ½′ in diameter; calyx-tube slightly enlarged, with spreading or incurved lobes; flesh thin and yellow; nutlets 2 or 3, slightly ridged on the back, about ¼′ long.

A tree, 20°—25° high, with a tall trunk 8′—10′ in diameter, covered with dark gray scaly bark, stout branches spreading into a broad flat-topped head, and slender branchlets covered at first with matted white hairs, becoming glabrous and light orange-brown at the end of their first season, and pale gray-brown the following year, and unarmed or armed with occasional slender nearly straight red-brown spines 1′—1½′ long.

Distribution. Borders of prairies and low moist soil a few miles west of Opelousas, St. Landry Parish, Louisiana.

22. Cratægus edura Beadl.

Leaves oblong-obovate, rounded and obtuse or occasionally acute at apex, gradually narrowed from above the middle to the cuneate base, and serrate only at the apex, nearly fully grown when the flowers open early in April and then thin, dark green and puberulous above especially on the midrib, very pale and villose below, and at maturity thick and coriaceous, 1¼′—1½′ long, and 1½′—1¾′ wide, with a slender midrib, and primary veins within the parenchyma; turning in the autumn orange, yellow, or brown; petioles slender, light yellow, pilose, ⅛′—¼′ in length. Flowers ⅜′—½′ in diameter, on short sparingly villose pedicels, in compact hairy 5—12-flowered corymbs; calyx narrowly obconic, glabrous or with a few hairs at the base, the lobes narrow, acuminate, glabrous; stamens 16—20; anthers pale yellow or nearly white; styles 2 or 3. Fruit ripening and falling in September, in few-fruited drooping clusters, subglobose, orange or yellow with a red cheek, about /16′ in diameter; calyx-lobes little enlarged, closely appressed, often deciduous; nutlets 2 or 3, rather obscurely ridged on the back, about ¼′ long.

A tree, 20°—25° high, with a trunk 6′—8′ in diameter, branches spreading out into a broad flat-topped head, and branchlets pilose when they first appear, soon glabrous, becoming reddish brown, unarmed or armed with chestnut-brown or gray spines 1½′—2′ long.

Distribution. Low woods near Opelousas, St. Landry Parish, Louisiana.

23. Cratægus crocina Beadl.

Leaves oblong-obovate, rounded or acute at apex, gradually narrowed and cuneate at the slender entire base, and sharply serrate above the middle with straight or incurved glandular teeth, when they unfold more or less pubescent, and at maturity subcoriaceous, dark green, lustrous and glabrous or glabrate above, pale and covered below with short matted pale hairs most abundant on the thin midrib and obscure primary veins, 1¼′—2′ long, and ½′—1′ wide; turning in the autumn orange, yellow, or brown; petioles slender, puberulous, about ¼′ in length; leaves at the end of vigorous shoots elliptic to oblong-obovate, acuminate more coarsely serrate, often 2½′ long and ⅚′ wide. Flowers opening at the end of April when the leaves are fully grown, ½′—⅝′ in diameter, on short villose pedicels, in compact few-flowered villose corymbs; calyx narrowly obconic, coated with matted white hairs, the lobes narrow, acute, entire or sparingly serrate, glabrous on the outer surface, slightly villose on the inner surface toward the apex; stamens 20; anthers yellow; styles usually 2 or 3. Fruit ripening in October, ellipsoidal or short-oblong, nearly ½′ long, yellow, the calyx prominent, with elongated mostly recurved lobes; nutlets usually 2, narrowed and acute at the ends, ridged on the back, about ¼′ long.

A tree, 18°—20° high, with a short trunk 4′—6′ in diameter, spreading branches forming a wide flat-topped head, and slender mostly unarmed branchlets covered at first with matted pale hairs, and dark orange-brown and puberulous in their first winter.

Distribution. Low woods near Opelousas, St. Landry Parish, Louisiana.

24. Cratægus fera Beadl.

Leaves oblong-obovate, rounded or rarely acute at apex, gradually narrowed and concave-cuneate at the slender entire base, and sharply serrate above the middle with straight or incurved teeth, fully grown when the flowers open the middle of April and then thin, covered above by short white hairs, and slightly villose along the midrib and veins below, and at maturity coriaceous, dark green, scabrate and lustrous on the upper surface, pale and puberulous on the lower surface on the slender midrib and obscure primary veins, 2½′—3′ long, and about ¾′ wide; turning in the autumn orange, yellow, or brown; petioles slender, pubescent early in the season, becoming puberulous, ⅜′—⅝′ in length; leaves at the end of vigorous shoots oblong-obovate, rounded or acute and often short-pointed at apex, coarsely serrate, often 2½′ long, and 1¼′ wide. Flowers: ½′ in diameter, on long slender villose pedicels, in broad lax compound many-flowered corymbs covered more or less thickly with white hairs; calyx-tube narrowly obconic, slightly hairy near the base, glabrous above, the lobes narrow, acuminate, entire or sparingly glandular-dentate, glabrous on the outer surface and puberulous on the inner surface; stamens 16—20; anthers light yellow; styles usually 2 or 3. Fruit ripening in September and October, on long slender pedicels, in few-fruited drooping clusters, globose or subglobose, bright red or scarlet, ⅜′ in diameter; flesh thin and mealy; calyx enlarged, with spreading or erect persistent lobes; nutlets 2 or 3, with a high narrow ridge, ¼′—/16′ long.

A tree, sometimes 20° high, with a trunk 8′—9′ in diameter, spreading branches forming a broad flat-topped head, and slender nearly straight branchlets, villose at first, becoming glabrous, pale reddish brown, ultimately ashy gray, and sometimes armed with slender straight spines 1′—1¼′ long.

Distribution. Low open Oak and Hickory woods near Opelousas, St. Landry Parish, and Natchitoches, Natchitoches Parish, Louisiana.

25. Cratægus Mohrii Beadl.

Leaves obovate or rhombic, acute or acuminate, gradually narrowed and cuneate at the entire base, and coarsely, occasionally doubly serrate above with straight or incurved teeth, when they unfold glabrous and slightly villose along the midrib and the lower side of the principal veins, nearly fully grown when the flowers open early in May, and at maturity thin and firm or subcoriaceous, dark green and very lustrous above, pale below, 1′—1½′ long, and ⅔′—1′ wide, usually with 4 pairs of thin primary veins, a stout midrib sometimes puberulous on the under side and bright red in the autumn; petioles frequently red at maturity, ⅓′—½′ in length; leaves at the end of vigorous shoots sometimes 3′ long and 2′ wide, mostly broad-elliptic, acute or acuminate, coarsely doubly serrate, and frequently divided toward the apex into short broad acute lobes; petioles, strait, glandular; petioles broadly winged, and occasionally glandular with minute dark glands. Flowers cup-shaped, about ¾′ in diameter, on slender elongated pedicels, in loose thin-branched many-flowered glabrous or villose corymbs; calyx-tube narrowly obconic, glabrous or occasionally pilose toward the base, the lobes linear-lanceolate, entire or finely glandular-serrate; stamens 20; anthers small, light yellow; styles 3—5, surrounded at base by a narrow ring of pale hairs. Fruit ripening about the middle of October, gracefully drooping on elongated thin bright red pedicels, in many-fruited clusters, subglobose to short-oblong, somewhat flattened at apex, full and rounded at base, bright orange-red, about ⅓′ in diameter; calyx prominent, with a short tube and usually erect lobes often deciduous before the fruit ripens; nutlets usually 3, about ¼′ long.

A tree, from 20°—30° high, with a tall straight trunk 6′—8′ in diameter, covered with thin ashy gray or light red-brown bark, sometimes armed with long slender or branched spines, spreading slightly pendulous branches forming a rather open broad symmetrical head, and branchlets furnished with thin nearly straight bright chestnut-brown shining spines 1′—1½′ long.

Distribution. Western Georgia to central Alabama and eastern Mississippi, and northward to middle Tennessee; abundant and of its largest size in the low flat woods near Birmingham, Jefferson County, Alabama, ascending into the poorer and drier soils of the neighboring hillsides and low mountain slopes.

II. Punctatæ.

Conspectus of the Arborescent Species.

Fruit usually short-oblong.

Anthers rose color or yellow; stamens 20; leaves obovate, often acutely lobed above the middle, their veins deeply impressed; fruit on stout pedicels, short-oblong to subglobose, flattened at the ends, dull red or bright yellow, marked by large pale dots.

26. C. punctata (A).

Anthers rose color; stamens 10—20; leaves oblong-obovate or oval, their veins not deeply impressed, fruit on long slender pedicels, short-oblong to obovoid, rounded at the ends, dull brick-red, marked by large pale dots.

27. C. pausiaca (A).

Fruit usually globose or subglobose.

Stamens 20.

Anthers pale yellow.

Corymbs villose.

Leaves obovate to oval or rarely rhombic, acute; fruit globose, or sometimes broader than high, dull red, marked by small pale dots.

28. C. collina (A, C).

Leaves obovate, oval, or ovate, acute or acuminate, incisely lobed; fruit globose, dull red.

29. C. amnicola (C).

Corymbs glabrous; leaves broadly oval to ovate, rounded or acute at apex, occasionally rounded at base, subcoriaceous; fruit subglobose to short-oblong, dull orange-red, marked by large pale dots.

30. C. fastosa (C).

Anthers rose color.

Leaves scabrate on the upper surface.

Leaves ovate, oval or rarely obovate, acuminate; flowers in compact usually 6—8-flowered corymbs.

31. C. silvestris (A).

Leaves obovate to rhombic, acute or rarely rounded at apex; flowers in wide usually 9—12-flowered corymbs.

32. C. verruculosa.

Leaves glabrous on the upper surface.

Corymbs slightly villose.

Leaves obovate to rhombic, acute or rounded at apex; fruit globose, dark dull red.

33. C. sordida (C).

Leaves oval to obovate, acute or acuminate at apex; fruit often rather longer than broad, bright canary-yellow.

34. C. brazoria (C).

Corymbs densely villose; leaves obovate, acute, acuminate or rounded at apex; fruit subglobose, dark dull red.

35. C. dallasiana (C).

Stamens 10.

Anthers pale yellow; leaves obovate, acute or acuminate or rounded and short-pointed at apex; fruit subglobose, pubescent at the ends, dull orange-red.

36. C. Lettermanii (A).

Anthers rose color; leaves oblong-obovate, acute or rounded at apex; fruit globose, bright scarlet, slightly pruinose.

37. C. pratensis (A).

26. Cratægus punctata Jacq.

Leaves obovate, pointed or rounded at apex, gradually narrowed to the cuneate entire base, sharply and often doubly serrate above the middle with minute teeth, and sometimes more or less incisely lobed, thickly covered below with pale hairs and pilose above when hey unfold, about half grown when the flowers open from the middle of May until early in June and then pilose on the midrib and veins below and nearly glabrous above, and at maturity thick and firm, pale gray-green and glabrous on the upper surface, more or less villose on the lower surface, 2′—3′ long, and ¾′—1½′ wide, with a broad prominent midrib, and primary veins deeply impressed on the upper surface; turning bright orange or orange and scarlet in the autumn; petioles stout, at first villose or tomentose, becoming pubescent or glabrous, ¼′—½′ in length; leaves at the end of vigorous shoots usually incisely lobed, and often 3′—4′ long and 1½′—2′ wide. Flowers ½′—¾′ in diameter, on slender villose pedicels, in tomentose or villose many-flowered compact corymbs; calyx-tube narrowly obconic, villose or tomentose, the lobes narrow, acute, nearly entire or minutely glandular-serrate, villose on the inner surface; stamens 20; anthers rose color or yellow; styles 5, surrounded at base by conspicuous tufts of white hairs. Fruit ripening and falling in October, on elongated nearly glabrous pedicels, in drooping clusters, short-oblong to subglobose, truncate at the ends dull red or bright yellow (var. aurea Ait.) and usually agreeing with the anthers in color, marked by numerous small white dots, ½′—1′ long; nutlets 5, about ¼′ long.

A tree, 20°—30° high, with a trunk occasionally a foot in diameter, stout branches spreading nearly at right angles and forming a round or flat-topped head, or sometimes ascending and forming a narrow open irregular head, and branchlets coated at first with pale deciduous pubescence, becoming light orange-brown or ashy gray, and armed with slender straight light orange-brown or gray spines 2′—3′ long.

Distribution. Common and generally distributed; rich hillsides; valley of the Chateaugay River, Quebec, to the valley of the Detroit River, Ontario, southward through western New England to Delaware, and along the Appalachian Mountains to northern Georgia, ascending in North Carolina and Tennessee to altitudes of nearly 6000°, and westward through New York, Ohio and Indiana to southern Michigan, Indiana, Illinois, southern Wisconsin, southeastern Minnesota, and in central Iowa. A form (var. canescens Britt.), densely hoary-tomentose on the under surface of the leaves, and on the petioles and corymbs, occurs in Bucks County, Pennsylvania, and near Albany, Albany County, New York; and a form (var. microphylla Sarg.) with smaller leaves and compact few-flowered corymbs has been found at Linesville, Crawford County, Pennsylvania.

27. Cratægus pausiaca Ashe.

Leaves oblong-obovate to oval, rounded or acute at apex, gradually narrowed from near the middle to the concave-cuneate entire base, and finely doubly serrate above with straight glandular teeth, more than half grown when the flowers open from the 20th to the end of May and then membranaceous, dark yellow-green, and slightly villose above and along the under side of the midrib and veins, and at maturity glabrous, dark yellow-green above, paler below, 2′—2½′ long, and 1¼′—1½′ wide, with a slender yellow midrib, and 5 or 6 pairs of primary veins extending very obliquely to the end of the leaf; petioles slender, wing-margined above the middle, villose only early in the season, ⅝′—1′ in length; leaves at the end of vigorous shoots elliptic to rhombic, long-pointed, slightly or deeply divided into broad lateral lobes, coarsely serrate, often 3½′—4′ long and 2′—2½′ wide. Flowers ½′ in diameter, on long slender hairy pedicels, in broad many-flowered thin-branched villose corymbs; calyx-tube narrowly obconic, villose below with closely appressed white hairs, glabrous above, the lobes abruptly narrowed from the base, slender, acuminate, tipped with minute dark glands, entire or occasionally obscurely toothed above the middle, glabrous on the outer surface, villose on the inner surface; stamens 10—15, rarely 20; anthers dark rose color; styles 2 or 3, surrounded at base by a broad ring of pale tomentum. Fruit ripening about the middle of October, on elongated slender slightly hairy pedicels, in drooping many-fruited clusters, short-oblong to obovoid, broad and rounded at the ends, dull brick-red, marked by large pale dots, /16′—/16′ long, and about ⅜′ thick; calyx small, with spreading appressed lobes mostly deciduous from the ripe fruit; flesh thin, hard, slightly juicy, green or greenish yellow; nutlets 3 or 4, thin, acute or obtuse at the ends, ridged on the back with a high broad deeply grooved ridge, about ¼′ long.

A tree, 20°—25° high, with a tall straight trunk often a foot in diameter, covered with dark brown scaly bark, stout wide-spreading branches forming a broad symmetrical round or flat-topped head, slender straight branchlets light orange-green and sparingly villose at first, becoming light orange-brown during their first season, light or dark gray-brown the following year, and armed with numerous stout slender straight orange-brown shining spines 1½′—2′ in length, long persistent on the branches and trunk, finally ashy gray, and becoming sometimes a foot long, with long slender lateral spines.

Distribution. Dry limestone hills and low moist bottom-lands, Bucks, Berks and Delaware counties, eastern Pennsylvania; at Chapin, Ontario County, New York.

28. Cratægus collina Chapm.

Leaves obovate to oval or occasionally to rhombic, acute, gradually narrowed or broadly cuneate at the entire base, and irregularly and often doubly serrate above with glandular incurved or straight teeth, when they unfold bright red and covered with soft pale hairs most abundant on the under side of the midrib and principal veins, less than one third grown when the flowers open at the end of April, and at maturity subcoriaceous, yellow-green on the upper surface, paler on the lower surface, glabrous with the exception of a few hairs on the under side of the stout yellow midrib and 4 or 5 pairs of slender primary veins, 1½′—2′ in length, and 1′—1¼′ wide; petioles slender, villose, soon glabrous, more or less winged toward the apex, ¼′—½′ long; leaves at the end of vigorous shoots frequently divided into short broad acute lateral lobes, more coarsely dentate and often 3′ long and 2½′ wide, with a stout broadly winged petiole generally light red like the lower side of the base of the midrib. Flowers ¾′ in diameter, on long stout pedicels, in broad many-flowered villose corymbs; calyx-tube broadly obconic, villose particularly toward the base, the lobes gradually narrowed from a broad base, acuminate, usually glabrous on the outer surface, villose on the inner surface, finely glandular-serrate with dark glands, bright red toward the apex; stamens usually 20; anthers large, pale yellow; styles 5. Fruit ripening in September, on stout elongated pedicels, in few-fruited erect or drooping puberulous clusters, subglobose but sometimes rather broader than long, dull red, marked by small pale dots, ⅓′—½′ in diameter; calyx enlarged, the lobes closely appressed, glandular-serrate, mostly persistent; flesh yellow; nutlets 5, broad and rounded at the ends, ridged and often grooved on the back, about ¼′ long.

A tree, usually 15°—20° but occasionally 25° high, with a tall straight trunk often buttressed at base, frequently armed with numerous large much-branched spines sometimes 6′—8′ long, stout wide-spreading branches forming a handsome flat-topped symmetrical head, and branchlets tinged with red and villose with long matted silky white hairs when they first appear, soon puberulous, and dull reddish brown, becoming gray in their second year, and furnished with stout lustrous spines 2′—3′ long.

Distribution. Hillsides in rich soil in the foothill region of the southern Appalachian Mountains from southwestern Virginia to central Georgia and westward to northeastern Mississippi and middle Tennessee; in central Alabama; ascending to altitudes of 2500° above the sea.

29. Cratægus amnicola Beadl.

Leaves obovate, oval or ovate, acute or acuminate at apex, gradually narrowed and concave-cuneate at the entire base, coarsely sometimes doubly serrate above with straight or incurved glandular teeth, and incisely lobed above the middle with short acute or acuminate lobes, deeply tinged with red and covered with short pale mostly caducous hairs when they unfold, about half grown and sparingly villose on the midrib and veins when the flowers open late in April or early in May, and at maturity subcoriaceous, bright green, glabrous, 1¼′—1½′ long, and 1′—1¼′ wide; turning in the autumn yellow, orange, red, and brown; petioles slender, sparingly villose early in the season, becoming glabrous, sometimes slightly glandular, ¼′—⅓′ in length; leaves at the end of vigorous shoots sometimes 2′ long and 1½′ wide. Flowers about ⅝′ in diameter, on elongated slender slightly villose pedicels, in narrow many-flowered villose corymbs; calyx-tube narrowly obconic, glabrous or with a few scattered hairs at the base, the lobes narrow, acuminate, glandular-serrate, glabrous; stamens 20; anthers nearly white; styles 3—5. Fruit on slender elongated glabrous pedicels, in drooping few-fruited clusters, subglobose, dull red, about ⅓′ in diameter; calyx enlarged, with elongated coarsely serrate reflexed conspicuous lobes; flesh yellow, thin, and firm; nutlets 3—5, rounded or slightly grooved on the back, nearly ¼′ long.

A tree, occasionally 25° high, with a trunk 8′—12′ in diameter, spreading or ascending branches forming a large wide head, and branchlets villose at first with long matted white hairs, soon glabrous, becoming orange-brown and ultimately ashy gray, and unarmed, or armed with stout spines 1¼′—2′ long.

Distribution. Low moist woods and the borders of streams, southeastern Tennessee, northwestern Georgia, and northeastern Alabama; common.

30. Cratægus fastosa Sarg.

Leaves broadly oval to ovate, rounded or acute at apex, concave-cuneate or rounded at the entire base, and coarsely doubly serrate above with straight glandular teeth, when they unfold covered above with long pale hairs and provided below with large tufts of snow-white tomentum in the axils of the primary veins, when the flowers open from the 20th to the 25th of April dark yellow-green and nearly glabrous on the upper surface and still tomentose in the axils of the veins below, and at maturity subcoriaceous, glabrous, yellow-green and lustrous above, pale yellow-green below, 1¾′—2′ long, and 1′—2′ wide, with a prominent light yellow midrib deeply impressed on the upper side, and usually 3—5 pairs of primary veins; petioles slender, at first densely villose, becoming puberulous, ½′—¾′ in length; leaves at the end of vigorous shoots occasionally lobed with broad acute lobes. Flowers about ¾′ in diameter, on slender pedicels, in compact many-flowered glabrous corymbs, with large conspicuous oblong-obovate and acute to lanceolate coarsely glandular-serrate bracts and bractlets usually persistent until after the petals fall; calyx broadly obconic, the lobes abruptly narrowed at base, slender, acuminate, coarsely glandular-serrate, glabrous on the outer surface, villose on the inner surface; stamens 20; anthers pale yellow; styles 5, surrounded at base by a broad ring of pale tomentum. Fruit ripening from the middle to the end of October, on thin reddish pedicels, in few-fruited drooping clusters, subglobose to short-oblong, dull orange-red, marked by large pale dots, ⅜′ in diameter; calyx enlarged, with spreading serrate lobes villose on the upper side, mostly deciduous from the ripe fruit; flesh thin, yellow-green; nutlets 3—5, thin, narrowed at the ends, obscurely ridged on the back with a broad low often grooved ridge, about /16′ long.

A tree, 18°—20° high, with a short trunk 8′—12′ in diameter, covered with dark brown or nearly black scaly bark, small ascending branches forming an irregular open head, and slender nearly straight branchlets, dark orange-green tinged with red when they first appear, becoming before autumn bright reddish brown and very lustrous, and dull reddish brown the following year, and armed with numerous stout nearly straight bright chestnut-brown shining spines 1½′—2′ long.

Distribution. Low woods near Fulton, Hemstead County, Arkansas; not common.

31. Cratægus silvestris Sarg.

Leaves ovate, oval or rarely obovate, acuminate, concave-cuneate or rounded at the entire base, sharply doubly serrate above with straight glandular teeth, and slightly divided above the middle into 3 or 4 pairs of small acuminate lobes, nearly fully grown when the flowers open at the end of May and then roughened above by short white hairs, and villose below on the slender midrib and veins, and at maturity subcoriaceous, dark yellow-green lustrous and scabrate on the upper surface, paler and still villose on the lower surface, 2¼′—2½′ long, and 1¾′—2′ wide; petioles stout, slightly hairy on the upper side, occasionally glandular, and ⅗′—⅘′ in length. Flowers ⅗′ in diameter, on slender villose pedicels, in compact villose usually 6—8-flowered corymbs; calyx-tube narrowly obconic, glabrous, the lobes slender, acuminate, coarsely glandular-serrate, slightly villose on the inner surface; stamens 20; anthers pink; styles usually 3. Fruit ripening at the end of September, on slender reddish slightly hairy pedicels, in few-fruited erect or spreading clusters, subglobose to short-oblong, truncate at base, rounded at apex, dull orange-red, about ½′ in diameter; calyx prominent with a broad deep cavity, and spreading coarsely serrate persistent lobes villose on the upper surface; flesh thick, dry and mealy; nutlets 3, gradually narrowed and rounded at the ends, ridged on the back with a high deeply grooved ridge, about ⅓′ long and ⅙′—⅕′ wide.

A tree, 30°—35° high, with a tall trunk often 1° in diameter, large ascending and spreading branches forming an open head, and stout nearly straight glabrous branchlets, light orange-green and marked by small pale lenticels when they first appear, becoming light chestnut-brown and lustrous in their first season, and dull red-brown the following year, and armed with slender straight or slightly curved dark chestnut-brown lustrous spines 1′—1½′ long.

Distribution. Woods in low moist soil, near London, Ontario.

32. Cratægus verruculosa Sarg.

Leaves obovate to rhombic, acute or rarely rounded at apex, cuneate and entire at base, and sharply often doubly serrate above with straight or incurved glandular teeth, when they unfold dark red, covered above by short pale hairs and below by long matted white hairs most abundant on the midrib and veins, about half grown when the flowers open from the 1st to the 10th of May and then thin, dark yellow-green and scabrate on the upper surface, and paler and pubescent on the lower surface, and at maturity subcoriaceous, dark green, lustrous and nearly smooth above, pale and still pubescent below on the stout midrib and conspicuous primary veins extending very obliquely toward the end of the leaf, 1½′—2′ long, and 1′—1¼′ wide; petioles stout, wing-margined at apex, at first villose, becoming pubescent or puberulous, ¼′—½′ in length; leaves at the end of vigorous shoots often broad-ovate to oval, sharply doubly serrate with straight teeth, sometimes slightly lobed above the middle with short acute lobes, and frequently 3′ long and 2′ wide. Flowers ¾′ in diameter, on long slender villose pedicels, in broad lax compound 6—12 usually 9-flowered villose corymbs; calyx-tube broadly obconic, thickly covered with matted pale hairs, the lobes gradually narrowed from a broad base, slender, acute, tinged with red at apex, sparingly glandular-serrate, pubescent; stamens 20; anthers pale rose color; styles 3—5 surrounded at base by a broad ring of long pale hairs. Fruit ripening about the 1st of October, on stout pubescent pedicels, in drooping few-fruited clusters, subglobose, somewhat flattened and pubescent at the ends, dark red; calyx prominent, with more or less deciduous lobes; nutlets 3—5, narrowed and acute at the ends, rounded and very irregularly ridged and sometimes obscurely grooved on the back, about ¼′ long.

A tree, 20°—25° high, with a tall trunk 10′—12′ in diameter, thick spreading branches forming a broad compact round-topped symmetrical head, and stout nearly straight branchlets thickly covered with matted pale hairs when they first appear, becoming reddish or orange-brown, nearly glabrous and roughened by minute tubercles at the end of their first season, gray-brown the following year, and armed with numerous straight stout or slender dark chestnut-brown very lustrous spines ¾′—1′ long.

Distribution. Springfield, Greene County, Missouri; not rare.

33. Cratægus sordida Sarg.

Leaves rhombic, acute, or occasionally obovate and rarely rounded at apex, cuneate and entire below, serrate above with narrow straight or incurved glandular teeth, and occasionally irregularly divided above the middle into short acute lobes, about half grown when the flowers open the first week of May and then membranaceous, bright green, lustrous and glabrous with the exception of a few short caducous hairs on the upper surface, particularly on the midrib and principal veins, and at maturity subcoriaceous, dark green and lustrous above, paler below, generally about 1½′ long and 1¼′ wide; petioles stout, slightly winged toward the apex, at first villose, soon glabrous, about ½′ long, often bright red in the autumn; leaves at the end of vigorous shoots sometimes oblong or oval, coarsely dentate, usually divided above the middle into short acute lobes, 3′—4′ long, 2′—2½′ wide, and decurrent on the stout glandular petioles. Flowers 1′—1¼′ in diameter, on slender pedicels, in few-flowered compact slightly villose corymbs; calyx-tube narrowly obconic, the lobes narrow, acuminate, villose on the inner surface; petals dull white; stamens 20; anthers small, rose color; styles 2 or 3, surrounded at base by a narrow ring of pale hairs. Fruit ripening and falling the middle of September, on short pedicels, in few-fruited drooping clusters, globose, ⅓′—½′ in diameter, dark dull red; calyx prominent, with elongated coarsely serrate appressed or incurved lobes; flesh thin and yellow; nutlets 2 or 3, broad, rounded and ridged on the back with a low rounded ridge, ¼′ long.

A slender tree, 20°—25° high, with a tall trunk 5′—6′ in diameter, often armed with long-branched spines, small ascending branches forming a narrow oval head, and slender nearly straight branchlets, dark orange-green and villose with long scattered pale hairs sometimes persistent until autumn, dull chestnut-brown in their second season, and dark gray-brown the following year, and furnished with numerous thin nearly straight bright chestnut-brown shining spines 1′—2½′ long, or often unarmed.

Distribution. Low woods and the gravelly banks of streams in Shannon, Carter, and Ripley Counties, southern Missouri.

34. Cratægus brazoria Sarg.

Leaves oval to obovate, acute or acuminate at apex, gradually narrowed, cuneate and entire at base, and coarsely and irregularly glandular-serrate above with straight spreading teeth, coated with hoary tomentum and often bright red when they unfold, nearly fully grown and covered with short soft pale hairs most abundant on the under side of the thin midrib and 3 or 4 pairs of primary veins when the flowers open from the middle to the end of March, and at maturity thin and firm in texture, glabrous, dark green and lustrous above, paler below, 2′—2½′ long, and 1¼′—1½′ wide; petioles slender, early in the season tomentose, becoming glabrous or puberulous, ½′—¾′ in length; leaves at the end of vigorous shoots broad-ovate or oblong, rounded or broad-cuneate at base, coarsely dentate, 5′ long, and 2½′ wide. Flowers ¾′ in diameter, on long slender pedicels, in broad slightly villose 7 or 8-flowered corymbs; calyx-tube narrowly obconic, coated with long matted pale hairs, the lobes narrow, acuminate, obscurely glandular-serrate or nearly entire, villose on both surfaces; stamens 20; anthers small, dark red; styles 5, surrounded at base by a thin ring of hoary tomentum. Fruit ripening after the 1st of October, in spreading or drooping few-fruited clusters, subglobose and often rather longer than broad, bright canary-yellow, marked by occasional dark dots, ⅓′—½′ long; calyx prominent, the lobes usually deciduous before the fruit ripens; flesh thin, light yellow, rather dry but sweet and edible; nutlets 5, rounded and grooved on the back, ¼′ long.

A tree, 20°—25° high, with a tall straight trunk 8′—10′ in diameter, numerous ascending branches forming a handsome symmetrical round-topped head, and branchlets covered when they first appear with matted pale hairs, dull reddish brown and often puberulous in their second season, and reddish brown the following year, and unarmed or occasionally armed with long thin gray spines.

Distribution. Low rich woods near the banks of the Brazos River, Columbia and Brazoria, Brazoria County, Texas.

35. Cratægus dallasiana Sarg.

Leaves oblong, acute, acuminate or rounded at apex, gradually narrowed to the concave-cuneate entire base, coarsely doubly serrate above with straight glandular teeth, and usually slightly lobed above the middle, coated below with thick hoary tomentum and villose above as they unfold, nearly fully grown and villose or tomentose below when the flowers open early in April, and at maturity thin, dark yellow-green, glabrous and lustrous on the upper surface, pale and pubescent on the lower surface on the slender midrib and 3 or 4 pairs of thin arching veins, 1¾′—2½′ long, and 1¼′—1½′ wide; petioles slender, wing-margined toward the apex, hoary-tomentose early in the season, becoming glabrous, about ½′ in length. Flowers about ⅝′ in diameter, on long slender hairy pedicels, in many-flowered densely villose corymbs; calyx-tube narrowly obconic, densely coated with long matted pale hairs, the lobes slender, acuminate, tipped with a minute red gland, sparingly and irregularly glandular-serrate, villose; stamens 20; anthers light rose color; styles 5. Fruit ripening at midsummer, on stout erect slightly hairy pedicels, in few-fruited clusters, subglobose, dull dark red, ⅜′—½′ in diameter; calyx prominent, with spreading lobes bright red on the upper side at the base; nutlets 5, acute at the narrow ends, thin, rounded and grooved with a broad shallow groove or irregularly ridged on the back, ¼′—/16′ long.

A tree, 20°—25° high, with a tall trunk 4′—6′ in diameter, covered with pale bark, small erect branches forming an open irregular head, and slender somewhat zigzag branchlets thickly coated at first with hoary tomentum, reddish brown and lustrous before autumn, ultimately ashy gray, and armed with straight slender gray-spines 1¼′—1½′ long.

Distribution. Forest-covered bottom-lands of the small tributaries of the Trinity River, Dallas County, Texas; not common.

36. Cratægus Lettermanii Sarg.

Leaves obovate, acute or acuminate or rounded and short-pointed at apex, gradually narrowed from near the middle and cuneate at the mostly entire base, coarsely often doubly serrate above with straight or incurved glandular teeth, and frequently slightly and irregularly divided above the middle into 3 or 4 pairs of short acute lobes, strongly plicate when they unfold and covered with a thick coat of pale tomentum, nearly half grown, roughened above by short pale hairs and pubescent below when the flowers open early in May, and at maturity thick, bright yellow-green and scabrate above, pale and pubescent below on the stout midrib and 4 or 5 pairs of primary veins, about 2′ long and 1½′ wide; petioles stout, more or less winged above the middle, at first tomentose, becoming pubescent or nearly glabrous, usually about ¾′ in length; leaves at the end of vigorous shoots broad-oval, acute or acuminate, more coarsely serrate, 2½′—3′ long, and 2′—2½′ wide. Flowers about ¾′ in diameter, on short villose pedicels in compact, many-flowered thick-branched densely villose corymbs; calyx-tube narrowly obconic, tomentose, the lobes narrow, acuminate, finely glandular-serrate, villose; stamens 10; anthers small, pale yellow; styles 5, surrounded at base by a broad ring of hoary tomentum. Fruit ripening early in October, on stout pubescent pedicels, in few-fruited spreading or drooping clusters, subglobose or occasionally slightly obovoid, rounded and puberulous at the ends, dull orange-red, marked by large pale dots, about ½′ in diameter; calyx broad, the lobes enlarged, coarsely glandular-serrate, reflexed, often deciduous before the fruit ripens; flesh thin; nutlets 5, prominently ridged on the back with a high rounded ridge, ¼′ long.

A tree, 18°—20° high, with a trunk 6′—8′ in diameter, with thin dark brown or nearly black bark separating freely into small plate-like scales, and often armed with thin much-branched spines frequently 7′—8′ long, small erect branches forming a wide open head, and branchlets coated when they first appear with hoary tomentum, dull red-brown, villose or pubescent during their first season, and furnished with stout straight bright red-brown shining spines 1½′—2′ long.

Distribution. Low rich soil inundated during several weeks in winter, among Oaks and Hickories; near Allenton, St. Louis County, Missouri.

37. Cratægus pratensis Sarg.

Leaves oblong-obovate, acute or rounded at apex, gradually narrowed below from near the middle to the cuneate entire base, sharply and often doubly serrate usually only above the middle with straight or incurved teeth tipped early in the season with a minute dark red caducous gland, and often more or less deeply divided toward the apex into short broad acute lobes, when they unfold bright bronze-yellow or dark red, and covered with short pale hairs, almost smooth and nearly fully grown when the flowers open at the end of May, and at maturity glabrous, thick, dark green and lustrous above, pale below, 1½′—2′ long, and 1′—1½′ wide, with a thin midrib, and 4 or 5 pairs of primary veins extending obliquely toward the end of the leaf, and raised and prominent below; petioles slender, glabrous, usually about ½′ in length; leaves at the end of vigorous shoots often oval or broad-ovate, frequently 3′ long and 2½′ wide. Flowers ⅓′ in diameter, on long slender pedicels, in broad loose many-flowered corymbs pubescent or puberulous at first but soon glabrous; calyx-tube narrowly obconic, coated toward the base with long matted pale hairs, the lobes narrow, acuminate, coarsely glandular-serrate, glabrous on the outer surface, villose on the inner surface; stamens 10; anthers small, rose color; styles 2 or 3, surrounded at base by a narrow ring of pale tomentum. Fruit ripening early in October and remaining on the branches until November, on elongated pedicels, in loose drooping many-fruited clusters, globose, bright scarlet, slightly pruinose, marked by occasional large pale dots, about ⅓′ in diameter; calyx prominent, with much enlarged coarsely glandular-serrate lobes often deciduous before the fruit becomes entirely ripe; flesh thin and yellow; nutlets 2 or 3, thick and broad, about ¼′ long.

A tree, occasionally 20° high, with a tall trunk 3′—7′ in diameter, often armed with long slender much-branched ashy gray spines, spreading branches forming a round-topped symmetrical head, and branchlets occasionally slightly villose when they first appear, soon glabrous, light orange-brown in their first season, and reddish or grayish brown the following year, and furnished with numerous thin straight or slightly curved shining chestnut-brown spines 2′—3′ long.

Distribution. Open woods near the banks of small streams in the prairie region of Stark and Peoria Counties, Illinois.

III. Æstivales.

Conspectus of the Arborescent Species.

Leaves glabrous with the exception of small axillary tufts of pale hairs on the lower surface, oblong-obovate; stamens 15—20; anthers pink or pale rose color.

38. C. æstivalis (C).

Leaves hoary-tomentose below early in the season, becoming villose with rufous hairs most abundant on the midrib and veins; stamens 20; anthers deep rose color.

Leaves oblong-obovate, acute or broad and rounded at apex, often slightly lobed above the middle, lustrous above; pedicels villose-pubescent.

39. C. rufula (C).

Leaves elliptic to oblong-cuneiform, narrowed at apex, dull above; pedicels glabrous.

40. C. opaca (C).

38. Cratægus æstivalis Sarg. May Haw. Apple Haw.

Mespilus æstivalis Walt.

Leaves oblong-obovate, rounded or acute at apex, gradually narrowed and cuneate at base, glabrous with the exception of small axillary tufts of pale hairs, and coarsely crenately serrate above the middle with gland-tipped teeth, beginning to unfold as the flowers open the middle of March, and when the fruit ripens at the end of May thin, dark green and lustrous above, yellow-green below, 1¼′—2′ long, and ⅓′—¾′ wide, with a slender yellow midrib and obscure primary veins; petioles slender, narrow wing-margined to below the middle, rarely furnished with occasional deciduous glands, about ¼′ in length; leaves at the ends of vigorous shoots elliptic to oblong-obovate, acute and usually abruptly short-pointed at apex, concave-cuneate at base, often lobed with one or two lateral lobes. Flowers ¾′ in diameter, on pedicels about ⅓′ long, in compact 2 or 3-flowered corymbs; calyx-tube narrowly obconic, glabrous, the lobes gradually narrowed from a broad base, short, entire, without glands, acute or acuminate and often red at apex, persistent and red on the fruit; stamens 15—20; anthers large, pink or pale rose color; styles usually 3. Fruit on a short slender erect pedicel, about ⅓′ long, usually solitary, short-oblong, scarlet, lustrous, about ⅓′ in length, the calyx persistent with erect lobes; flesh yellow, juicy, acidulous; nutlets usually 3, acute at ends, rounded and slightly ridged on the back, ¼′ long.

A slender tree, 20°—25° high, with a tall stem 6′—8′ in diameter, covered with pale flaky bark, erect or slightly spreading branches forming a narrow head, and slender straight or slightly zigzag branchlets chestnut-brown and lustrous during their first season, and dull gray-brown the following year, and armed with stout straight gray spines ½′—1¼′ in length.

Distribution. Low river banks, the borders of swamps and in depressions filled with water during most of the year; banks of the Ogeechee River near Meldrim, Effingham County, and near Valdosta, Lowndes County, Georgia; swamp of the Combahee River near Yemassee, Hampton County, and near Aiken, Aiken County, South Carolina; pond holes eight or nine miles west of Newbern, Craven County, North Carolina; passing into var. maloides Sarg. with young leaves tinged with red and villose along the upper side of the midrib, those at the end of vigorous shoots sometimes broad-obovate, rounded and divided at apex into 3 short rounded lobes, longer acuminate calyx-lobes and dark red anthers. Wet prairies, Volusia County, Florida; and into var. cerasoides Sarg. differing in the presence of short white hairs on the upper surface of the young leaves, in the longer acuminate calyx-lobes slightly villose on the inner surface and often minutely serrate near the middle, in the dark rose-colored anthers, and the late ripening fruit up to ½′ in diameter, on drooping pedicels often ½′ in length. An arborescent shrub with a round-topped head 30°—40° across, numerous large erect and spreading stems often 30° high, covered with smooth pale bark separating into thin plate-like scales, in falling disclosing the dull red inner bark, and slender nearly straight glabrous branchlets armed with straight slender spines 1′—1½′ in length. Fruit ripening late in July and in August. Low, wet, often inundated prairies near Sewall, Valusia County, Bradfordville, Leon County, Jasper, Hamilton County, and Quincy, Gadsden County, Florida. A form of this variety growing in Valusia County (f. luculenta Sarg.) differs in the more numerous hairs on the upper surface of the young leaves, in the rather smaller flowers, smaller and less juicy fruit ripening at the end of June or early in July, and in its often arborescent habit.

39. Cratægus rufula Sarg.

Cratægus æstivalis Torr. & Gray in part, not Mespilus æstivalis Walt.

Leaves oblong-obovate, acute or rounded at apex, gradually narrowed, cuneate and entire at base, finely crenately glandular-serrate, and often slightly lobed above the middle; with short rounded lobes, covered above with soft pale hairs and whitish tomentose below when they unfold, and at maturity thick, dark green, lustrous and glabrous or slightly pubescent along the midrib on the upper surface, rufous-pubescent especially on the midrib and veins on the lower surface, 1½′—2′ long, and ¾′—1¼′ wide, rarely not more than 1′ long and ½′ wide; petioles slender, villose-pubescent with rufous hairs, occasionally glandular, ¼′—⅓′ in length; leaves at the ends of vigorous shoots oblong-obovate, rounded and short-pointed to elliptic and acuminate, laterally lobed, or deeply 3-lobed at apex, often 2½′ long and 1½′ wide. Flowers appearing from the 10th to the end of March, ¾′—1′ in diameter, in mostly 3—5-flowered clusters, on villose-pubescent pedicels about ⅓′ in length; calyx-tube broadly obconic, glabrous or villose-pubescent sometimes in the same cluster, the lobes gradually narrowed from a broad base, acuminate, entire or slightly glandular-serrate nearly to apex, glabrous or slightly pubescent on the outer surface; stamens 20; anthers dark rose color; styles 3—5, surrounded at base by a ring of white tomentum. Fruit ripening at the end of May, often solitary on glabrous erect pedicels ¼′—½′ long, subglobose, scarlet, lustrous, about ½′ in diameter, the calyx persistent with erect lobes; nutlets only slightly grooved on the back, about ¼′ long.

A tree, sometimes 30° high, with a tall trunk 8′—10′ in diameter, covered with rough deeply furrowed dark bark, paler and less deeply furrowed on smaller and younger stems, stout ascending and spreading branches forming a broad round-topped head, and slender slightly zigzag branchlets covered when they first appear with pale tomentum, glabrous or rusty tomentose until the early summer, becoming chestnut-brown, lustrous and glabrous before autumn and dull gray in their second year, and unarmed or armed with slender or stout straight spines ½′—1½′ in length.

Distribution. Depressions filled with water except at midsummer, sandy borders of ponds and streams and low wet prairies, Cottondale and Round Lake, Jackson County, and Quincy, Gadsden County, Florida; near Bainbridge, Decatur County, and Albany, Dougherty County, Georgia; near Dothan, Houston County, Alabama; pond holes along the Neuse River near Goldsboro, Wayne County, North Carolina.

40. Cratægus opaca Hook.

Cratægus æstivalis Torr. & Gray in part, not Mespilus æstivalis Walt.

Leaves elliptic to oblong-cuneiform, gradually narrowed and acute or bluntly pointed at apex, cuneate at the often glandular base, finely crenately serrate above the middle with minute glandular teeth, pilose above and hoary-tomentose below when they unfold, and at maturity dull dark green and glabrous or slightly hairy on the midrib on the upper surface, pubescent on the lower surface with rusty brown hairs most abundant on the midrib and veins, 2′—2½′ long, and ½′—1′ wide; petioles slender, villose-pubescent, about ¼′ in length; leaves at the end of vigorous shoots elliptic to oblong-ovate, often irregularly laterally lobed, and 2½′—3′ long and wide. Flowers appearing in February and March before or with the unfolding of the leaves, 1′ in diameter, on glabrous pedicels ½′ long, in 3—5-flowered corymbs; calyx-tube broadly obconic, glabrous, the lobes narrowed from a wide base, short, nearly triangular, acute and tipped at apex with a conspicuous gland, entire or minutely serrate, glabrous, often deeply tinged with red; stamens 20; anthers large, deep rose color; styles 3—5, surrounded at base by a broad ring of pale tomentum. Fruit ripening early in May, in usually 2—3-fruited clusters, depressed-globose, scarlet, lustrous, dotted with pale spots, ½′—⅔′ in diameter, with a small narrow cavity surrounded by the erect calyx-lobes; nutlets 3—5, rounded at the ends, rounded and slightly grooved on the back, ¼′ long.

A tree, 20°—30° high, with a tall stem occasionally 1° in diameter, covered with deeply fissured bark, divided into dark red-brown persistent scales, slender mostly erect branches forming a narrow round-topped head, and slender branchlets villose-pubescent when they first appear, soon glabrous, lustrous and bright chestnut-brown during their first season, becoming dull gray in their second year, and armed with stout straight chestnut-brown spines ½′—1′ in length, or more often unarmed; occasionally with several stems forming a large shrub.

Distribution. In deep depressions filled with water for most of the year, low river banks and borders of swamps; near Mt. Vernon, Mobile County, and near Selma, Dallas County, Alabama; southern Mississippi (Meridian, Lauderdale County, and Hattiesburg, Forest County); eastern Louisiana; sometimes in St. Tammany Parish covering large tracts almost to the exclusion of other plants; western Louisiana from the coast to nearly the northern border of the state, and eastern Texas to the valley of the Trinity River; rare and local east of the Mississippi River; common westward. The fruit is largely used in making preserves and jellies.

IV. Virides.

Conspectus of the Arborescent Species.

Stamens 20.

Fruit not exceeding ⅓′ in diameter.

Anthers pale yellow.

Corymbs, branchlets and leaves glabrous.

Bark of the trunk pale gray, close and smooth.

Leaves ovate to oblong-obovate, acute or acuminate, rarely rounded at apex; fruit depressed-globose, bright scarlet or orange.

41. C. viridis (A, C).

Leaves ovate, acute, often broadly cuneate at base; fruit subglobose, orange-red.

42. C. ovata (A).

Leaves oval or ovate, acute, rounded or broadly cuneate at base; fruit globose, yellow-green flushed with red.

43. C. vulsa (C).

Bark of the trunk dark brown or nearly black; leaves subcoriaceous.

Leaves oblong-ovate to semiorbicular, acute, often short-pointed or rarely rounded at apex; fruit short-oblong to obovoid or globose, dull orange color.

44. C. glabriuscula.

Leaves oval to rhombic, acute or acuminate; fruit subglobose to short-oblong, bright orange-red.

45. C. blanda (C).

Corymbs and branchlets villose-pubescent; leaves ovate or obovate, acute or rounded at apex; fruit subglobose, orange-red.

46. C. velutina (C).

Anthers deep rose color; leaves elliptic to oblong-ovate, acute, acuminate or rarely rounded at apex; fruit globose or subglobose, orange-red.

47. C. arborescens (C).

Fruit ½′—¾′ in diameter.

Anthers yellow.

Leaves cuneate at base; calyx-tube glabrous.

Leaves lanceolate to oblong-obovate, acuminate; fruit short-oblong, dull brick red covered with a glaucous bloom.

48. C. nitida (A).

Leaves obovate to oval or rhombic, acute or rarely rounded at apex; fruit subglobose to short-oblong, dark crimson.

49. C. mitis (A).

Leaves, broad and rounded at base, ovate, acute; calyx-tube villose; fruit subglobose to short-oblong, dark red.

50. C. atrorubens (A).

Anthers rose color; corymbs villose; fruit red.

Leaves obovate, oval or ovate, acute, scabrate above; fruit globose to subglobose, anthers deep rose color.

51. C. ingens (C).

Leaves broadly obovate, oval or ovate, acute or acuminate, smooth above; fruit globose or depressed-globose; anthers pale rose color.

52. C. penita (C).

Stamens usually 10; occasionally 12—20; anthers bright red; leaves oblong-obovate to oval, usually acute or acuminate; fruit subglobose to short-oblong, bright orange-red.

53. C. micracantha (C).

41. Cratægus viridis L.

Cratægus Davisii Sarg.

Leaves ovate to oblong-obovate or oval, acute or acuminate or rarely rounded at apex, gradually narrowed to the cuneate base, finely serrate above with incurved glandular teeth, and sometimes slightly 3-lobed toward the apex, tinged with red and slightly hairy above when they unfold, nearly fully grown when the flowers open in April and May, and at maturity membranaceous to subcoriaceous, dark green and lustrous on the upper surface, paler on the lower surface, with large axillary tufts of pale hairs, 1′—2′ long, and ½′—1′ wide, with a thick midrib and conspicuous primary veins; often turning brilliant scarlet late in the autumn before falling; petioles slender, 1′—1½′ in length; leaves at the end of vigorous shoots often deeply laterally lobed with narrow acuminate lobes, and 2½′—4′ long, and 1½′—2′ wide. Flowers ¾′ in diameter, on long slender pedicels, in many-flowered corymbs; calyx-tube narrowly obconic, glabrous, the lobes lanceolate, entire; stamens 20; anthers pale yellow; styles 2—5, usually 5, surrounded at base by conspicuous tufts of pale hairs. Fruit ripening in the autumn and mostly persistent on the branches through the winter, on long slender pedicels, in drooping many-fruited clusters, depressed-globose, bright scarlet or orange, ⅛′—¼′ in diameter; calyx little enlarged, the lobes often deciduous from the ripe fruit; nutlets usually 5, narrowed and rounded at the ends, rounded and slightly grooved or ridged on the back, /16′—⅛′ long.

A tree, 20°—35° high, with a straight often fluted trunk 8°—12° tall, and 18′—20′ in diameter, covered with close gray or pale orange-colored bark, small branches forming a round rather compact head, and slender glabrous branchlets ashy gray to light red-brown in their first winter, and unarmed or occasionally armed with slender sharp pale spines ¾′—1′ long.

Distribution. On the often inundated borders of streams and swamps, rarely in drier ground on low slopes; southeastern Virginia (banks of the Blackwater River near Zuni, Isle of Wight County), North Carolina (Salisbury, Rowan County), South Carolina (near Aiken, Aiken County), eastern Georgia (near Augusta, Richmond County, and Macon, Bibb County), western Florida (River Junction, Gadsden County, and Tallahassee, Leon County to the swamps of the lower Apalachicola River), and westward through central and southern Alabama, southern Mississippi, and Louisiana to the valley of the San Antonio River (Sutherland Springs, Wilson County), Texas, and to central and western Arkansas, eastern Oklahoma and southeastern Missouri (Butler County), and northward in the region adjacent to the Mississippi River from Louisiana to northeastern Missouri, and to Pike County, Illinois, ranging eastward in Mississippi to Tishomingo County in the northeastern corner of the state, to northwestern Georgia, southeastern Tennessee, and to Richland County, Illinois; rare and local in the Atlantic and east Gulf states; common and often forming great thickets in western Louisiana, the coast region of eastern Texas, southern Arkansas, and in the region adjacent to the Mississippi River.

42. Cratægus ovata Sarg.

Leaves ovate, acute, broadly or acutely concave-cuneate at the entire base, coarsely often doubly serrate above with glandular teeth, and occasionally slightly divided into short lateral lobes, nearly fully grown when the flowers open early in May and then dark green, very smooth and glabrous above with the exception of a few short scattered hairs near the base of the midrib, paler below, with small persistent axillary tufts of white hairs, and at maturity membranaceous, 2′—2½′ long, and 1½′—2′ wide, with a slender yellow midrib and primary veins; petioles slender, rose-colored in the autumn, about ¾′ in length; leaves at the end of vigorous shoots rounded or truncate at the broad base, coarsely serrate, and sometimes 3′ long and wide. Flowers about ½′ in diameter, on long slender pedicels, in broad loose many-flowered corymbs; calyx-tube narrowly obconic, glabrous, the lobes broad acute, entire or coarsely glandular-serrate toward the apex, glabrous; styles 5. Fruit ripening in October, on elongated pedicels, in long drooping clusters, subglobose or a little longer than broad, orange-red, ¼′—/16′ long; calyx enlarged, with elongated closely appressed lobes sometimes deciduous from the ripe fruit; nutlets 5, acute at the ends, rounded or slightly ridged on the back, about /16′ long.

A tree, 25°—30° high, with a tall trunk sometimes a foot in diameter, covered with smooth gray bark, slender glabrous branchlets light reddish brown and lustrous during their first year, becoming grayish brown in their second season, and unarmed or armed with occasional dark purple slender slightly curved shining spines 1′ long.

Distribution. Low moist soil on the banks of the River Desperes, South St. Louis, St. Louis County, and near Alba, Jasper County, Missouri.

43. Cratægus vulsa Beadl.

Leaves oval or ovate, acute, broad and rounded or broad-cuneate at the entire base, irregularly and often doubly serrate above with straight or incurved gland-tipped teeth, and often divided into several short acute lateral lobes, when they unfold dark bronze-red, and pilose with scattered caducous hairs, and furnished below with tufts of pale often persistent hairs in the axils of the principal veins, nearly fully grown when the flowers open late in April, and at maturity thin, bright green on the upper surface, paler on the lower surface, about 2′ long and 1½′ wide, with a slender midrib and 4 or 5 pairs of thin pale yellow primary veins; turning in the autumn yellow or brown; petioles slender, somewhat villose at first, soon becoming glabrous, about ¾′ in length; leaves at the end of vigorous shoots broadly ovate, acute or acuminate, broad and rounded or occasionally truncate or broadly cuneate at base, more coarsely serrate and more deeply lobed, often 3′ long and 2½′ wide, with a stout winged glandular petiole. Flowers ¾′ in diameter, on slender pedicels, in compact 3—10-flowered corymbs, with linear acuminate glandular red bracts and bractlets; calyx-tube broadly obconic, glabrous, the lobes gradually narrowed from a broad base, acuminate, entire or occasionally obscurely serrate toward the apex, glabrous; stamens 20; anthers pale yellow; styles 3—5, surrounded at base by a thin ring of pale hairs. Fruit ripening at the end of September or early in October, on slender pedicels, in few-fruited drooping clusters, globose, yellow-green flushed with red, ⅓′ in diameter; calyx prominent, with closely appressed lobes; flesh yellow-green; nutlets 3—5, thin, rounded, sometimes slightly ridged and grooved on the back, about /16′ long.

A tree, occasionally 20° high, with a tall trunk 5′—6′ in diameter, covered with thin fissured bark separating into light gray scales tinged with brown, and often armed with long compound spines, ascending or spreading branches forming an oval usually compact symmetrical head, and slender nearly straight glabrous chestnut-brown branchlets becoming gray, and armed with thin nearly straight bright chestnut-brown shining spines 1′—1½′ long; sometimes a shrub, with numerous stems.

Distribution. Rich moist soil in the neighborhood of streams; northwestern Georgia and northeastern Alabama.

44. Cratægus glabriuscula Sarg.

Leaves oblong-ovate to semiorbicular, acute or often short-pointed or rarely rounded at apex, gradually narrowed from below the middle to the slender entire base, coarsely and often doubly serrate usually only above the middle with broad straight gland-tipped teeth, and sometimes divided toward the apex into 2 or 3 short acute lobes, nearly fully grown when the flowers open the 1st of April, and then membranaceous and slightly pilose above with scattered hairs most abundant along the base of the midrib, and at maturity subcoriaceous, hard and firm, dark green and lustrous on the upper surface, pale on the lower surface, 1½′—2′ long, and ¾′—1′ wide, with a thin light yellow midrib, and primary veins extending obliquely toward the end of the leaf, conspicuous secondary veins and reticulate veinlets; petioles slender, wing-margined, ⅓′ in length; leaves at the end of vigorous shoots often ovate, broadly cuneate at base, much more coarsely serrate, more frequently lobed, 2′—2½′ long and wide. Flowers about ½′ in diameter, on long slender pedicels, in few-flowered rather compact corymbs; calyx-tube broadly obconic, glabrous, the lobes short, gradually narrowed from a broad base, entire, villose on the inner surface; stamens 20, anthers nearly white; styles 5. Fruit ripening in September and often persistent until late into the winter, on long slender pedicels, in compact many-fruited drooping clusters, short-oblong to obovoid or nearly globose, dull orange color, marked by minute dark dots, about ¼′ long; calyx enlarged, conspicuous, with spreading or closely appressed lobes dull red on the upper side at base, often deciduous before the fruit ripens; flesh very thin, yellow, dry and hard; nutlets 5, rounded and sometimes obscurely grooved on the back, about /16′ long.

A tree, 20°—25° high, with a tall straight trunk often a foot in diameter, covered with thin dark brown scaly bark, long ascending branches forming a narrow head, and slender nearly straight branchlets, unarmed or armed with occasional slender straight chestnut-brown lustrous spines ¾′—1′ long.

Distribution. Bottom-lands of the Trinity River and its branches near Dallas, Dallas County, Texas, in forests of Elms and Nettle-trees.

45. Cratægus blanda Sarg.

Leaves oval to rhombic, acute or acuminate, and occasionally slightly lobed toward the apex, broadly cuneate or concave-cuneate at the entire base, coarsely crenately serrate above the middle with gland-tipped teeth, coated with soft pale hairs when they unfold, fully grown when the flowers open about the 1st of May, and then membranaceous, dark green and lustrous above and glabrous below with the exception of large axillary tufts of snow-white tomentum, and at maturity subcoriaceous, yellow-green and lustrous on the upper surface, paler on the lower surface, 1½′—2′ long, and 1′—1⅓′ wide, with a slender midrib, and 2 or 3 pairs of thin primary veins extending obliquely toward the end of the leaf; petioles slender, at first villose along the upper side, soon becoming glabrous, ¾′—1′ in length; leaves at the end of vigorous shoots often broadly ovate, rounded at base, more deeply lobed above the middle, 2′—2½′ long, and 1½′—2′ wide. Flowers 1′ in diameter, on slender elongated pedicels, in broad many-flowered corymbs, with linear entire bracts and bractlets; calyx-tube broadly obconic, glabrous, the lobes gradually narrowed from a broad base, acuminate, entire or obscurely dentate, glabrous; stamens 20; anthers canary-yellow; styles 5. Fruit ripening about the middle of October, on slender pedicels, in many-fruited drooping clusters, subglobose to short-oblong, bright orange-red, ¼′ in diameter; calyx prominent, with spreading lobes usually deciduous from the ripe fruit; nutlets 5, thin, narrowed at the ends, deeply grooved on the back, ¼′ long.

An unarmed tree, 25°—30° high, with a tall trunk 10′—12′ in diameter, covered with dark brown or nearly black bark divided by shallow fissures and broken on the surface into small plate-like scales, stout ascending branches forming a broad irregular head, and nearly straight glabrous branchlets dark orange-green at first, becoming dull red-brown during their first season and darker brown the following year.

Distribution. Dry uplands and low rolling hills near Fulton, Hempstead County, Arkansas.

46. Cratægus velutina Sarg.

Leaves ovate to obovate, acute or rounded at apex, gradually narrowed and cuneate at the entire base, and sharply often doubly serrate with straight glandular teeth, more than half grown when the flowers open at the end of April and then covered above by short white hairs and below with hoary pubescence, and often furnished with axillary tufts of white tomentum, and at maturity glabrous, smooth and lustrous on the upper surface and covered on the lower surface with matted pale hairs, 1¾′—2′ long, and 1½′—2′ wide, with a thin midrib and primary veins; petioles slender, thickly covered early in the season with matted hairs, becoming glabrous, ½′—1′ in length; leaves at the end of vigorous shoots ovate, rounded or broad-cuneate at base, coarsely serrate, usually slightly lobed above the middle, and often 2½′—3′ long and 1½′ wide. Flowers ½′ in diameter on slender villose pedicels, in usually 7—12-flowered hairy corymbs; calyx-tube narrowly obconic, villose, the lobes gradually narrowed from a broad base, short, acute, entire, slightly villose; stamens 20; anthers yellow; styles 5. Fruit on long slender glabrous or nearly glabrous drooping stems in few-fruited clusters, subglobose, orange-red, marked by small pale dots, about ¼′ in diameter; calyx prominent, with a deep narrow cavity pointed in the bottom, and closely appressed lobes; flesh thin, dry and mealy; nutlets 5, acute at base, rounded at apex, ridged on the back with a low grooved ridge, about ⅙′ long and ⅛′ wide.

A tree, 20°—25° high, with a trunk 8′—10′ in diameter, covered with dark rough scaly bark, and slender slightly zigzag branchlets, hoary-tomentose when they first appear, light reddish brown, marked by pale lenticels and glabrous or sometimes pubescent near the end in their first autumn, and ashy gray the following year, and armed with slender nearly straight chestnut-brown spines 1′—1½′ in length.

Distribution. Uplands in dry sandy soil, Fulton, Hempstead County, near Texarkana, Bowie County, Arkansas; and in the valley of the lower Brazos River (near Columbia, Brazoria County), Texas.

47. Cratægus arborescens Ell.

Leaves elliptic to oblong-obovate, acute, acuminate or rarely rounded and abruptly short-pointed and slightly lobed at apex, gradually narrowed cuneate and entire at base, and coarsely doubly serrate above the middle with incurved glandular teeth, villose on the upper side of the midrib with short white hairs when they unfold, and at maturity thin, glabrous, dark green and lustrous on the upper surface, paler and often furnished on the lower surface with small axillary tufts of pale hairs, 1′—2′ long, and ¾′—1′ wide, with a slender midrib and primary veins; petioles slender, glabrous, ½′—1′ in length; leaves at the end of vigorous shoots oval to oblong-ovate or elliptic, acuminate, abruptly or gradually narrowed and cuneate at base, more or less deeply lobed with acuminate lateral lobes, often 2½′ long and 1¼′ wide, their petioles stout, and glabrous early in the season. Flowers ½′ in diameter, on slender pedicels, in wide many-flowered compound corymbs; calyx-tube narrowly obconic, glabrous or slightly pilose, the lobes slender, acuminate, entire, glabrous or slightly villose on the inner surface, deciduous from the ripe fruit; stamens 20; anthers deep rose color; styles usually 5. Fruit on short pedicels in many-fruited drooping clusters, globose or subglobose, orange-red, ¼′—⅓′ in diameter; nutlets 5, pointed at the ends, slightly ridged on the back, about ⅙′ long.

A tree, 25°—30° high, with a tall trunk 12′—18′ in diameter covered with close pale gray bark, spreading and erect branches forming a broad rather open irregular head, and slender glabrous red-brown branchlets, ashy gray in their second season, and unarmed or armed with straight slender chestnut-brown spines.

Distribution. River banks, low wet woods and borders of swamps; Georgia-coast region, near Dorchester, Liberty County, in the neighborhood of Savannah, and on the Ogeechee River at Fort Argyle, Chatham County (type station); near Augusta, Richmond County, Georgia.

48. Cratægus nitida Sarg.

Leaves lanceolate to oblong-obovate, acuminate, abruptly or gradually narrowed and cuneate at the entire base, coarsely serrate above with straight or incurved glandular teeth, and often more or less divided into 2 or 3 pairs of broad acute lobes, dark red and slightly villose along the upper side of the midrib with scattered caducous hairs when they unfold, nearly fully grown when the flowers open early in May, and at maturity thick and coriaceous, dark green and very lustrous on the upper surface, pale and dull on the lower surface, 2′—3′ long, and 1′—1½′ wide, with a prominent midrib usually red on the lower side, and few thin prominent primary veins generally extending to the point of the lobes; turning in the autumn rich orange color through shades of bronze and orange-red; petioles stout, glandular, villose while young on the upper side, soon becoming glabrous, ½′—¾′ in length; leaves at the end of vigorous shoots more deeply lobed and frequently 5′ long and 2½′ wide. Flowers ¾′ in diameter, on long slender pedicels, in broad compound many-flowered glabrous corymbs; calyx-tube narrowly obconic, glabrous, the lobes slender, elongated, acuminate, entire or sparingly glandular-serrate; stamens 15—20; anthers pale yellow; styles 2—5. Fruit ripening at the end of October, on slender elongated pedicels, in many-fruited drooping clusters, short-oblong, full and rounded at the ends, pruinose with a glaucous bloom, marked by small dark dots, ½′—⅝′ long, and about ⅓′ in diameter; calyx only slightly enlarged, the lobes dark red at the base on the upper side, usually erect, often deciduous before the fruit ripens; nutlets 2—5, rounded and ridged on the back with a broad low rounded ridge, light-colored, ¼′ long.

A tree, often 30° high, with a tall straight trunk sometimes 18′ in diameter, covered with close dark bark broken into thick plate-like scales, stout spreading lower branches and erect upper branches forming a broad often irregular head, and slender glabrous branchlets bright orange-brown and lustrous during their first and second seasons, becoming pale reddish brown in their third year, and ultimately ashy gray, and unarmed or armed with occasional straight thin bright chestnut-brown lustrous spines 1′—1½′ long.

Distribution. Bottoms of the Mississippi River, St. Clair County, Illinois; common.

49. Cratægus mitis Sarg.

Leaves obovate to oval or rhombic, acute or rarely rounded at apex, gradually narrowed and concave-cuneate at the entire base, and coarsely serrate above with straight glandular teeth, nearly fully grown when the flowers open during the first week of May, and then light yellow-green above, paler below, and glabrous with the exception of a few short hairs on the upper side of the midrib, and at maturity subcoriaceous, dark green and lustrous on the upper surface, pale yellow-green on the lower surface, 1½′—2½′ long, and 1′—1½′ wide, with a prominent midrib and slender primary veins; petioles stout, wing-margined at apex, occasionally glandular with minute glands, 1⅓′—1½′ in length. Flowers ½′—⅝′ in diameter, on long slender pedicels, in compact 8—15-flowered glabrous corymbs, with red glandular bracts and bractlets; calyx-tube narrowly obconic, glabrous, the lobes glabrous, abruptly narrowed from a broad base, acuminate, finely glandular-serrate below the middle with minute stipitate red glands; stamens 20; anthers yellow; styles 2—4, usually 3. Fruit ripening the middle of October, on slender pedicels, in many-fruited drooping clusters, subglobose to short-oblong, rounded at the ends, dark crimson, marked by occasional large dark dots, ½′—⅝′ long, about ½′ in diameter; calyx only slightly enlarged, the lobes serrate, closely appressed, often deciduous from the ripe fruit; flesh thick, pale orange color, and juicy; nutlets usually 3, thick, full and rounded at the ends, prominently ridged on the back, with a broad high rounded deeply grooved ridge, about ¼′ long.

A tree, 25°—30° high, with a tall trunk sometimes a foot in diameter, covered with dark scaly bark, large spreading branches forming a broad round-topped head, and glabrous branchlets dull light reddish brown during their first season, becoming dark brown or ashy gray, and armed with stout straight or slightly curved dull red-brown or purplish spines usually about 1½′ long.

Distribution. Low moist rich soil on the bottoms of the Mississippi River near the village of Kahokia, St. Clair County, Illinois.

50. Cratægus atrorubens Ashe.

Leaves ovate, acute, usually rounded or sometimes cuneate or truncate at the broad entire base, coarsely and usually doubly serrate above, and often divided into 2 or 3 pairs of short acute lobes, about half grown when the flowers open late in April or early in May and then slightly roughened above by short scattered white hairs, and furnished below with conspicuous axillary tufts of pale tomentum, and at maturity thin, glabrous, dark dull green and smooth on the upper surface, light yellow-green on the lower surface, about 2′ long and 1½′ wide, or on vigorous shoots frequently 3′ long, and 2½′ wide, with a thin midrib and 4 or 5 pairs of slender primary veins; petioles slender, nearly terete, more or less densely villose early in the season, soon becoming glabrous, 1′—1½′ in length. Flowers about ⅝′ in diameter, on slender elongated villose pedicels, in broad loose glabrous or villose corymbs; calyx-tube narrowly obconic, densely villose throughout or only at base with pale tomentum, the lobes short, acute, finely glandular-serrate, villose particularly on the inner surface; stamens 20; styles 4 or 5, surrounded at base by a narrow ring of pale tomentum. Fruit ripening and falling early in October, on slender pedicels, in drooping few-fruited clusters, subglobose to short-oblong, rounded at the ends, dark red; calyx somewhat enlarged, with spreading lobes usually deciduous before the fruit ripens; nutlets 4 or 5, thin, rounded and sometimes obscurely grooved on the back, about /16′ long.

A tree, sometimes 30° high, with a tall trunk 12′—18′ in diameter, covered with dark red-brown scaly bark, thin erect and spreading branches forming a compact rather narrow head, and slender glabrous branchlets marked by occasional dark lenticels, dark green more or less tinged with red when they first appear, soon becoming dark chestnut-brown and very lustrous, and bright reddish brown in their second year, and usually unarmed.

Distribution. St. Louis County, Missouri, and rich bottom-lands of the Mississippi River, St. Clair County, Illinois; not common.

51. Cratægus ingens Beadl.

Leaves obovate-oval or ovate, broadly or acutely cuneate at the entire base, crenately serrate above, and often slightly lobed toward the acute apex, about half grown when the flowers open at the end of April or early in May and then roughened above by short rigid hairs and villose below along the midrib, and the remote slender veins extending obliquely to the point of the lobes, and at maturity subcoriaceous, dark green and scabrate on the upper surface, paler and nearly glabrous on the lower surface, 1½′—2′ long, and 1¼′—1½′ wide; turning in the autumn yellow, orange, red, or brown; petioles stout, narrowly wing-margined to the middle, pubescent while young, becoming glabrous, about ¾′ in length; leaves at the end of vigorous shoots more deeply lobed and often 3′—3½′ long, and 2′ wide, with a stout broad-winged petiole sometimes 1½′ long. Flowers ½′—⅝′ in diameter, on slender hairy pedicels, in many-flowered compact hairy corymbs; calyx narrowly obconic, coated, especially toward the base with matted pale hairs, the lobes slender, elongated, acute, glandular with bright red glands, glabrous on the outer, sparingly villose on the inner surface; stamens 20; anthers deep rose color; styles 3—5. Fruit ripening in October, on stout puberulous pedicels, in few-fruited drooping clusters, globose to subglobose, red, about ⅜′ in diameter; calyx little enlarged, with reflexed appressed nearly glabrous lobes; nutlets 3—5, rounded or slightly grooved and ridged on the back, ¼′ long.

A tree, sometimes 25° high, with a trunk a foot in diameter, spreading branches forming a wide round-topped head, and unarmed branchlets covered at first with matted pale hairs, soon becoming glabrous, dark chestnut-brown.

Distribution. Moist woods and the low banks of streams; southeastern Tennessee and northwestern Georgia.

52. Cratægus penita Beadl.

Leaves broad-obovate, oval, or ovate, acute or acuminate at apex, broadly or acutely concave-cuneate at the entire base, sharply often doubly serrate above with glandular mostly straight teeth, and often slightly lobed above the middle, deeply tinged with red and covered with pale hairs when they unfold, nearly fully grown when the flowers open about the 1st of May and then smooth above, and glabrous below with the exception of axillary tufts of pale hairs, and at maturity subcoriaceous, dark green and lustrous on the upper surface, paler on the lower surface, 1¾′—2′ long, and 1′—1¾′ wide, with a prominent midrib and slender primary veins; turning orange, yellow, and brown in the autumn; petioles slender, covered while young like the upper side of the base of the midrib with pale deciduous hairs, ½′—¾′ in length; leaves at the end of vigorous shoots often rounded or subcordate at base, more or less deeply lobed, and 2½′—3′ long and broad, with a stout broadly winged glandular petiole. Flowers about ⅝′ in diameter, on elongated glabrous or sparingly hairy pedicels, in compact few-flowered nearly glabrous corymbs; calyx broadly obconic, glabrous, the lobes gradually narrowed from a broad base, slender, acuminate, entire, or furnished with occasional minute glandular teeth, slightly villose on the inner surface; stamens 20; anthers white faintly tinged with pink; styles 3—5. Fruit ripening in October, on elongated slender pedicels, in few-fruited drooping clusters, globose or depressed-globose, red, about ¼′ in diameter; calyx enlarged, with spreading or reflexed lobes villose on the upper side; nutlets 3—5, narrowed and acute at the ends, rounded and broadly grooved on the back, about ¼′ long.

A tree, 18°—20° high, with a short trunk sometimes 10′ in diameter, stout ascending or spreading branches forming a wide head, unarmed branchlets puberulous while young, soon glabrous, becoming light reddish brown.

Distribution. Low moist woods and the banks of streams; southeastern Tennessee.

53. Cratægus micracantha Sarg.

Leaves oblong-obovate to oval, acute, acuminate, or rarely rounded at apex, gradually or abruptly narrowed from above or from below the middle to the cuneate entire base, coarsely crenulate-serrate, and occasionally 3-lobed above with short broad acute lateral lobes, when they unfold villose on the upper and hoary-tomentose on the lower surface, more than half grown when the flowers open about the middle of May and then membranaceous and slightly villose above with short scattered pale hairs, and at maturity thin but firm in texture, dark yellow-green, lustrous and smooth above, paler and tomentose below on the slender midrib and 3 or 4 pairs of very obscure primary veins, 2′—2½′ long, and 1′—1¼′ wide; petioles slender, tomentose early in the season, becoming glabrous or pubescent, ½′—1′ in length; leaves at the end of vigorous shoots often broadly rhombic to obovate, acuminate, frequently deeply 3-lobed or divided into 2 or 3 pairs of short lateral lobes, usually 2½′—3′ long. Flowers cup-shaped, ¼′ in diameter, on long slender pedicels thickly coated with matted white hairs, in broad lax many-flowered compound hairy corymbs; calyx-tube narrowly obconic, villose, the lobes linear, acuminate, entire, slightly villose, tipped with minute dark glands; stamens usually 10, occasionally 12, 15, or 20; anthers small, deep bright red; styles 5. Fruit ripening the middle of October, on slender pubescent pedicels, in drooping many-fruited clusters, subglobose to short-oblong, full and rounded at the ends, bright orange-red, lustrous, marked by occasional large pale dots, about ¼′ long; calyx prominent, with a short villose tube, and spreading erect hairy lobes often deciduous from the ripe fruit; nutlets 5, thin, acute at the narrow ends, rounded and sometimes slightly grooved on the back, about /16′ long.

An unarmed tree, sometimes 25° high, with a tall trunk 8′—12′ in diameter, covered with light or dark brown bark separating freely into thin narrow scales, stout spreading branches forming a broad flat-topped handsome head, and slender nearly straight branchlets coated until after the flowering time with thick hoary tomentum, bright red-brown and puberulous during their first season, becoming light or dark dull reddish brown the following year.

Distribution. Common in low woods in rich moist soil near Fulton, Hempstead County, Arkansas.

V. Pruinosæ.

Conspectus of the Arborescent Species.

Stamens 20.

Anthers rose color.

Leaves elliptic; fruit subglobose, green and pruinose when fully grown, becoming dark purple-red and very lustrous; anthers large, deep rose color.

54. C. pruinosa (A, C).

Leaves ovate, acute or acuminate; fruit short-oblong, dull russet-green; anthers small, light rose color.

55. C. georgiana (C).

Anthers white; leaves ovate, acute, cordate at base; fruit broader than high, scarlet, pruinose, becoming lustrous.

56. C. callicarpa (A).

Stamens 10; anthers dark rose color; leaves broad-ovate, acuminate; fruit subglobose, green more or less tinged with red, pruinose.

57. C. disjuncta (A).

54. Cratægus pruinosa K. Koch.

Leaves elliptic, acute, broadly or acutely cuneate at the entire base, irregularly and often doubly serrate above with glandular straight or incurved teeth, and divided in 3 or 4 pairs of short acute or acuminate lateral lobes, when they unfold bright red and glabrous with the exception of a few short caducous hairs on the upper side of the base of the midrib, nearly fully grown when the flowers open from the middle to the end of May and then membranaceous and bluish green, and at maturity subcoriaceous, dark blue-green and often glaucous above, pale below, 1′—1½′ long, and ¾′—1′ wide, with a slender midrib, and 3 or 4 pairs of thin primary veins running to the point of the lobes; late in the autumn turning dull orange color; petioles slender, glandular, slightly winged at the apex, often bright red in early spring and in the autumn, 1′—1½′ in length; leaves at the end of vigorous shoots broad-ovate, often rounded at base, more coarsely serrate and more deeply lobed, frequently 2½′ long and wide, with stouter and more broadly winged petioles. Flowers ¾′—1′ in diameter, on long slender pedicels, in few-flowered glabrous corymbs; calyx-tube broadly obconic, glabrous, the lobes gradually narrowed from a wide base, long-pointed, finely glandular-serrate below the middle; stamens 20; anthers large, deep rose color; styles 5, surrounded at base by a thick ring of hoary tomentum. Fruit on long thin light green ultimately bright red pedicels, in few-fruited drooping clusters, 5-angled, apple green and covered with a glaucous bloom until nearly fully ripe, at maturity late in October subglobose but rather broader than high, barely angled, ½′—⅝′ in diameter, dark purple-red, marked by many small dull dots, very lustrous; calyx prominent, with a long well-developed tube, and enlarged usually erect lobes often deciduous before the fruit ripens; flesh thick, light yellow; nutlets 5, light-colored, acute at apex, narrowed and rounded at base, deeply grooved on the back, ¼′ long.

A tree, 15°—20° high, with a stem a few inches in diameter, spreading horizontal branches forming a broad open irregular head, and slender glabrous branchlets bright chestnut-brown during their first season, later becoming dark reddish brown, and armed with numerous stout straight light chestnut-brown spines 1′—1½′ long; often shrubby, with several intricately branched stems.

Distribution. Slopes of low hills often in limestone soil; southwestern Vermont, westward through New York to southern Ontario (neighborhood of Toronto), and through Ohio and Indiana to central and northern Illinois, and southward through eastern Pennsylvania to northern Delaware.

55. Cratægus georgiana Sarg.

Leaves ovate, acute or acuminate at apex, rounded or broad-cuneate at base, finely and often doubly serrate with straight or incurved gland-tipped teeth, and divided into numerous short acute lateral lobes, glabrous with the exception of a few pale caducous hairs on the upper surface and bronze-yellow when they unfold, nearly half grown when the flowers open about the 20th of April and then thin, dark yellow-green above and pale below, and at maturity thin but firm in texture, dark blue-green on the upper surface, pale on the lower surface, 1½′—2′ long, and 1′—1¼′ wide, with a slender yellow midrib and 3 or 4 pairs of thin primary veins; petioles slender, often short-winged at the apex, usually about ¾′ in length; leaves at the end of vigorous shoots often 3′ long and 2′ wide, sometimes deltoid and usually much more deeply lobed. Flowers ¾′ in diameter, on slender pedicels, in usually 5—7-flowered compact glabrous corymbs; calyx-tube broadly obconic, glabrous, the lobes gradually narrowed from a broad base, acuminate, entire or obscurely and irregularly serrate, glabrous; stamens 20; anthers small; light rose color; styles 5; surrounded at the base by a narrow ring of pale tomentum. Fruit ripening and falling early in October, on slender pedicels, in drooping few-fruited clusters, short-oblong, full and rounded at the ends, often obscurely 5-angled, dull russet-green, ⅜′—½′ long; calyx-lobes only slightly enlarged, mostly deciduous before the fruit ripens, leaving a well-defined ring at the summit of the short calyx-tube; flesh thin, light green; nutlets 5, thin, rounded and irregularly grooved on the back, about ¼′ long.

A tree, sometimes 25°—30° high, with a tall trunk 10′—12′ in diameter, stout wide-spreading branches forming a broad symmetrical round-topped head, and slender lustrous chestnut-brown branchlets armed with straight or slightly curved thin spines rarely more than 1½′ long.

Distribution. Low rich river-bottoms and meadows in the neighborhood of Rome, Floyd County, Georgia.

56. Cratægus callicarpa Sarg.

Leaves ovate, acute, cordate at base, coarsely often doubly serrate with long straight glandular teeth, and slightly divided into 3 or 4 pairs of short broad acuminate lateral lobes, not more than a quarter grown when the flowers open late in April and then very thin, yellow-green and slightly villose above and on the midrib below, and at maturity thin, glabrous, dark yellow-green and lustrous on the upper surface, pale yellow-green on the lower surface, 4′—4½′ long, and 2′—2½′ wide, with a stout midrib, and 3 or 4 pairs of prominent primary veins connected by conspicuous cross veinlets; petioles stout, slightly wing-margined at apex, sparingly glandular, 1′—1¼′ in length; leaves at the end of vigorous shoots thicker, with shorter glandular petioles rose-colored toward the base. Flowers 1′ in diameter, on short stout pedicels, in small compact 5—10-flowered corymbs, with lanceolate to linear-obovate glandular bracts and bractlets usually persistent until the flowers open; calyx-tube broadly obconic, glabrous, the lobes separated by wide sinuses, short, broad, acuminate, coarsely glandular-serrate, slightly villose on the inner surface; stamens 20; anthers white; styles 5, surrounded at base by a broad ring of pale tomentum. Fruit ripening early in October on short stout spreading pedicels in 2 or 3-fruited clusters, broader than high, distinctly 5-angled, rounded at the wide apex, truncate at base, with a deep depression at the insertion of the pedicel, scarlet, pruinose, becoming lustrous, marked by numerous large pale dots, ¾′—⅘′ broad, and about ¾′ high; calyx-lobes deciduous; flesh thin, light yellow slightly tinged with red, remaining on the ground through the winter without becoming soft; nutlets 5, thin, acute at apex, rounded at base, rounded and slightly grooved or ridged with a low grooved ridge on the back, ⅕′—¼′ long and wide.

A tree, 20°—25° high, with a tall stem 5′—6′ in diameter covered with dark scaly bark, and stout nearly straight branchlets dark orange-green when they first appear, becoming light chestnut-brown, lustrous and marked by small pale lenticels in their first season, and dull reddish brown the following year, and armed with stout straight or slightly curved purplish spines 1′—1½′ in length.

Distribution. Rich hillsides, near Shrewsbury, St. Louis County, Missouri.

57. Cratægus disjuncta Sarg.

Leaves broad-ovate, acuminate, full and rounded or concave cuneate at the entire base, sharply often doubly serrate above with straight or incurved glandular teeth, and slightly and irregularly divided above the middle into narrow acuminate spreading lobes, thin, glabrous, dark blue-green above, pale below, 2½′—3′ long, and 2¼′—2½′ wide, with a slender yellow midrib, and 4 or 5 pairs of thin primary veins extending obliquely to the point of the lobes; petioles slender, wing-margined at apex, glandular, 1′—1¼′ in length. Flowers opening the first of May, ⅔′ in diameter, on long stout pedicels, in glabrous compact 3—6 usually 5-flowered glabrous corymbs, with conspicuous glandular early deciduous bracts and bractlets; calyx-tube narrowly obconic, glabrous, the lobes slender, acuminate, glabrous, entire or sparingly glandular-serrate; stamens 10; anthers large, dark rose color; styles 4 or 5, surrounded at base by a narrow ring of pale tomentum. Fruit on stout rigid pedicels, in drooping or spreading clusters, subglobose, usually rather broader than high, angled, green more or less tinged with red, pruinose, ½′—¾′ in diameter; calyx prominent, with a short tube and much enlarged spreading or erect lobes usually deciduous at midsummer; flesh thin, greenish yellow; nutlets usually 4, rounded at the ends, deeply grooved on the back, about ¼′ long.

A tree, 15°—18° high, with a tall slender trunk, covered with dark slightly scaly bark, small erect and spreading branches forming an open irregular head, and stout slightly zigzag glabrous branchlets dark olive-green tinged with red when they first appear, dark dull reddish brown or purple and marked by small pale lenticels at the end of their first season, becoming light grayish brown in their second year, and armed with numerous stout nearly straight dark purple lustrous spines 2½′—3′ in length.

Distribution. Gravelly banks of small streams near Monteer, Shannon County, and at Carl Junction, Jasper County, Missouri.

VI. Silvicolæ.

Medioximæ Sarg.

Conspectus of the Arborescent Species.

Fruit on short erect pedicels; leaves rounded or occasionally slightly cordate at base.

58. C. drymophila (C).

Fruit on elongated drooping pedicels.

Leaves truncate, cordate or rounded at base; anthers, pale rose.

59. C. diffusa (A).

Leaves cuneate or rounded at base; anthers, dark purple.

60. C. luxuriosa (A).

58. Cratægus drymophila Sarg.

Cratægus silvicola Beadl.

Leaves ovate, acute or acuminate at apex, rounded at the entire base, sharply and often doubly serrate above with gland-tipped teeth, and slightly and irregularly divided into short acute lateral lobes, when they unfold dark red and coated with short soft pale hairs most abundant on the upper surface, about half grown when the flowers open at the end of April and then nearly glabrous, and at maturity thin, dark yellow-green and smooth or scabrate above, pale and glabrous below, or occasionally villose along the under side of the slender midrib, and of 3 or 4 pairs of thin primary veins extending to the point of the lobes, about 2′ long and 1½′—1¾′ wide; petioles slender, glandular, about 1′ in length; leaves at the end of vigorous shoots often deltoid, truncate or cordate at base, more coarsely serrate, more deeply lobed, and often 2½′ long and wide. Flowers about ¾′ in diameter, on slender pedicels, in compact few-flowered thin-branched glabrous corymbs, with linear glandular bright red caducous bracts and bractlets; calyx-tube narrowly obconic, glabrous, the lobes gradually narrowed, acuminate, glabrous, entire or glandular-serrate; stamens 10; anthers large, dark rose color; styles 3—5, surrounded at base by a narrow ring of pale hairs. Fruit ripening at the end of September and soon falling, on short pedicels, in erect few-fruited clusters, subglobose and often a little broader than long, red or greenish yellow, with a rosy cheek, about ½′ in diameter; calyx little enlarged, with spreading lobes usually deciduous before the fruit ripens; flesh thin and yellow; nutlets 3—5, about ¼′ long.

A tree, sometimes 30° high, with a tall straight trunk 6′—8′ in diameter, covered with close or slightly fissured bark broken into small gray or red-brown scales, and often armed with long stout branched gray spines, ascending or spreading branches forming a narrow irregular or round-topped head, and slender branchlets dark green tinged with red and covered with long pale scattered white hairs when they first appear, soon becoming glabrous, bright red-brown during their first year, and ultimately ashy gray, and armed with few or many thin straight or somewhat curved bright chestnut-brown spines 1½′—2′ long; or in dry soil of upland forests usually a shrub, with numerous stems.

Distribution. Low moist flat woods; northern Alabama and northwestern and central Georgia, and occasionally on the drier uplands of the surrounding country; common; central Mississippi (Pelahatchee, Rankin County; Jackson, Hinds County, and in Franklin County); eastern Louisiana (Holtsville, St. Tammany Parish, anthers pink, R. S. Cocks).

59. Cratægus diffusa Sarg.

Cratægus Beckwithæ Sarg.
Cratægus Robbinsiana Sarg.

Leaves ovate, acute or acuminate at apex, rounded, truncate or cordate at the entire base, often doubly serrate above with straight glandular teeth, and more or less deeply divided into 4 or 5 pairs of spreading acuminate lateral lobes, deeply-tinged with red, glabrous below and covered above with short white hairs when they unfold, nearly fully grown when the flowers open from the middle to the 20th of May and then thin, pale yellow-green and hairy above and pale below, and at maturity thin and firm, smooth, dark green and glabrous on the upper surface, pale on the lower surface, 1¼′—2′ long, and 1′—1½′ wide, with a slender yellow midrib, and thin primary veins extending obliquely to the point of the lobes; often turning orange color tinged with red in the autumn; petioles slender, slightly wing-margined at apex, glandular with minute stipitate dark glands, ½′—¾′ in length; leaves at the end of vigorous shoots broad-ovate, usually long-pointed, cordate or rarely truncate at base, more coarsely serrate, more deeply lobed, and frequently 2½′—3′ long, and 2′—2½′ wide, with a stout reddish conspicuously glandular petiole ⅖′—⅗′ in length. Flowers ½′—⅗′ in diameter, on slender glabrous pedicels, in 6—10-flowered corymbs, with linear glandular bracts and bractlets mostly deciduous before the flowers open; calyx-tube broadly obconic, glabrous, the lobes gradually narrowed from a wide base, acuminate at the gland-tipped apex, entire or slightly and irregularly toothed near the middle; stamens 7—10; anthers light rose color; styles 4 or 5, surrounded at base by a ring of pale tomentum. Fruit ripening from the first to the middle of October, on slender pedicels, in few-fruited erect clusters, depressed-globose, rather broader than high, dull red and slightly pruinose, becoming lustrous, and about ½′ in diameter; calyx little enlarged, with spreading appressed lobes bright red on the upper side below the middle and mostly persistent on the ripe fruit; flesh thin, hard, greenish white; nutlets 4 or 5, broad and rounded at base, narrowed and rounded at apex, ridged on the back with a high ridge, about ¼′ long.

A tree, occasionally 30° high with a tall trunk 8′—10′ in diameter, covered with light gray closely appressed scales, comparatively small erect branches forming an open head, and slender slightly zigzag branchlets marked by numerous dark lenticels, green tinged with red and glabrous when they first appear, bright chestnut-brown and lustrous during their first winter, and pale gray-brown the following year, and armed with numerous slender or occasionally stout nearly straight bright red-brown shining spines 1¼′—1½′ long; usually smaller and sometimes a shrub.

Distribution. Valley of the Connecticut River (Walpole, Cheshire County, New Hampshire, and Westminster and Putney, Windham County, Vermont), western Vermont (near Burlington, Chittenden County); eastern, central and western New York; common.

60. Cratægus luxuriosa Sarg.

Leaves oblong-ovate, acuminate, gradually narrowed and cuneate or rounded at the often unsymmetrical base, finely often doubly serrate with straight glandular teeth, and slightly divided usually only above the middle into 3 or 4 pairs of small acute lobes, about half grown when the flowers open late in May and then thin, dark yellow-green and roughened above by short white hairs and paler below, and at maturity thin, dark yellow-green and scabrate on the upper surface, pale bluish green on the lower surface, 2¼′—2½′ long, and 1¾′—2′ wide, with a slender midrib and obscure primary veins; petioles slender, slightly wing-margined at apex, occasionally glandular with minute persistent glands, 1′—1¼′ in length; leaves at the end of vigorous shoots broad-ovate, rounded at base, coarsely serrate, laterally lobed with numerous short broad lobes, often 3′ long and 2½′ wide. Flowers ¾′ in diameter, on short slender pedicels, in compact mostly 6—12-flowered corymbs; calyx-tube narrowly obconic, the lobes long, slender, acuminate, entire or occasionally slightly dentate near the middle, glabrous on the outer surface, slightly villose on the inner surface; stamens 8—10; anthers bright purple; styles 3—5. Fruit ripening and beginning to fall early in October, on short stout pedicels, in drooping usually 1—3-fruited clusters, subglobose to slightly obovoid, scarlet, lustrous, marked by pale dots, ½′—¾′ in diameter; calyx little enlarged, with a deep narrow cavity and spreading and incurved usually persistent lobes dark red on the upper side below the middle; flesh thick, yellow-green and acid; nutlets 3—5, usually 4, gradually narrowed and rounded at the ends, ridged on the back with a broad high grooved ridge, about ¼′ long.

An oval-headed tree, 20°—30° high, with a short trunk sometimes 8′—10′ in diameter, covered with dark gray scaly bark, and stout zigzag often contorted branchlets dark orange-green and marked by large pale lenticels when they first appear, becoming light chestnut-brown and lustrous in their first season and dull red-brown the following year, and armed with few stout slightly curved chestnut-brown shining spines 1′—1½′ long, persistent and becoming branched on old stems.

Distribution. Rich hillsides, Kittanning, Armstrong County, and on the flood plain of the Allegheny River at Whiskey Hollow across the river from Kittanning, and Linesville, Crawford County, Pennsylvania.

VII. Tenuifoliæ.

Conspectus of the Arborescent Species.

Stamens 5—10.

Corymbs villose.

Leaves oblong-ovate; stamens usually 5; anthers pink; fruit obovoid to short-oblong.

61. C. apiomorpha (A).

Leaves oblong-obovate; stamens 10; anthers reddish purple; fruit obovoid to subglobose.

62. C. paucispina (A).

Corymbs glabrous; leaves oval or ovate; stamens usually 5; anthers dark reddish purple; fruit short-oblong.

63. C. pentandra (A).

Stamens usually 20.

Corymbs villose.

Leaves broad-ovate to obovate or rarely oval; fruit short-oblong to obovoid.

64. C. lucorum (A).

Leaves rhombic to broad-ovate or rarely obovate; fruit ellipsoidal.

65. C. lacera (C).

Corymbs glabrous.

Leaves ovate; anthers pale rose color; fruit subglobose to broad-obovoid, dark red.

66. C. depilis (A).

Leaves ovate; stamens 15—20; anthers dark rose color; fruit subglobose.

67. C. basilica (A).

61. Cratægus apiomorpha Sarg.

Leaves oblong-ovate, acuminate, rounded or rarely cuneate at the entire often unsymmetrical base, finely doubly serrate above with slender glandular teeth, and slightly divided above the middle into 4 or 5 pairs of triangular acute lobes, about half grown when the flowers open early in May and then membranaceous, light yellow-green and tinged with red or bronze color, and covered above with short white hairs and pale and glabrous below, and at maturity thick and firm in texture, dark blue-green and smooth and lustrous or sometimes dull and scabrate on the upper surface, pale blue-green on the lower surface, 1½′—2¼′ long, and 1⅛′—1½′ wide, with a stout midrib, and primary veins arching obliquely to the point of the lobes; petioles slender, slightly winged at the apex, often sparingly glandular, ⅜′—1′ in length; leaves at the end of vigorous shoots often 3′ long. Flowers ½′—¾′ in diameter, on short villose or glabrous pedicels, in compact many-flowered usually hairy corymbs, their bracts and bractlets linear to oblong-obovate, glandular-serrate with stipitate dark red or purple glands, turning red before falling, mostly persistent until after the flowers open; calyx-tube narrowly obconic, glabrous, the lobes abruptly narrowed at base, slender, acuminate, entire or sparingly glandular; stamens 5—10, usually 5; anthers pink; styles 3—5, surrounded at base by tufts of pale hairs. Fruit ripening early in September and soon falling, on slender pedicels, in few-fruited drooping clusters, obovoid or rarely short-oblong, bright reddish purple, marked by small scattered pale dots, ⅜′—⅝′ long, and ¼′—⅓′ in diameter; calyx much enlarged, with spreading lobes, their tips mostly deciduous from the ripe fruit; flesh thin, yellow, juicy, pleasantly acid; nutlets 3—5, thin, rounded and ridged on the back with a low ridge, about ¼′ long.

A tree, sometimes 25° high, with a trunk 6′ in diameter and 3°—6° long, covered with dark gray bark separating into thin plates, in falling disclosing the yellow inner bark, numerous ascending branches forming an oblong or pyramidal crown, and slender branchlets dark dull red-brown during their first season, becoming dark gray-brown the following year, and unarmed, or armed with slender nearly straight dull red-brown ultimately ashy gray spines 1′—1½′ long; or often shrubby, with numerous stems spreading into small clumps.

Distribution. Dry open places, borders of woods, and the margins of the high banks of streams; common and generally distributed in northeastern Illinois.

62. Cratægus paucispina Sarg.

Leaves oblong-obovate, acuminate, rounded, concave-cuneate to truncate or subcordate at the entire base, sharply doubly serrate above with straight glandular teeth, and deeply divided into 4 or 5 pairs of acute lateral lobes spreading or pointing toward the apex of the leaf, about half grown when the flowers open early in May and then light yellow-green and slightly roughened above by short white hairs and paler and glabrous below, and at maturity membranaceous, dark blue-green and scabrate on the upper surface, pale blue-green on the lower surface, 2½′—3′ long, and 1½′—2½′ wide, with a slender yellow midrib, and thin primary veins extending obliquely to the point of the lobes; petioles slender, usually without glands, tinged with purple in the autumn, ¾′—1½′ in length. Flowers ⅝′—¾′ in diameter, on slender hairy pedicels, in broad 12—20-flowered slightly villose corymbs, their bracts and bractlets linear to oblong-obovate, glandular, red, mostly persistent until after the flowers open; calyx-tube narrowly obconic, glabrous, the lobes narrow, acuminate, glandular-serrate with small dark red stipitate glands, glabrous on the outer, pubescent on the inner surface; stamens 10; anthers bright reddish purple; styles 4 or 5, surrounded at base by tufts of pale hairs. Fruit ripening during the first half of September and soon falling, on slender glabrous pedicels, in drooping clusters, obovoid to subglobose, crimson or purplish, marked by numerous small pale dots, slightly pruinose, ½′—⅝′ long, and about ½′ in diameter; calyx small, with reflexed and appressed or erect and incurved serrate lobes dark red on the upper side below the middle, often deciduous from the ripe fruit; flesh thin, yellow, juicy, acid and edible; nutlets 4 or 5, thin, narrowed and acute at the ends, rounded and slightly grooved or obscurely ridged on the back, about ¼′ long.

A tree, sometimes 25° high, with a trunk 4′—6′ in diameter and often 6° long, covered with dark gray or nearly black bark separating into thin plate-like scales, numerous branches forming a round-topped head, and slender glabrous branchlets dark yellow-green when they first appear, becoming dark reddish brown at the end of their first season, olive-green in their second year, and ultimately dark gray-brown, and armed with small straight light red-brown shining spines ½′—¾′ long.

Distribution. Woods and river banks in dry clay soil; northeastern Illinois; common.

63. Cratægus pentandra Sarg.

Leaves oval or ovate, acuminate, broadly cuneate or rarely rounded at the entire base, divided above the middle into numerous short acute or acuminate lobes, and coarsely and often doubly serrate with straight or incurved teeth tipped with small dark glands, nearly fully grown and very thin when the flowers open at the end of May, and at maturity membranaceous, dark green and roughened above by short rigid pale hairs, pale and glabrous below, 2′—2½′ long, and 1½′—2′ wide, with a slender yellow midrib, and thin primary veins extending to the point of the lobes; petioles slender, often winged toward the apex, glandular with minute dark glands, usually about 1′ in length; leaves at the end of vigorous shoots more deeply lobed, and often 4′ long and 3′ wide. Flowers ⅝′—¾′ in diameter, on long slender pedicels, in compact few-flowered glabrous corymbs; calyx-tube narrowly obconic, glabrous, dark red, the lobes linear-lanceolate, entire or finely glandular-serrate; stamens usually 5, occasionally 6—10; anthers large, dark red-purple; styles 3, surrounded at base by a thin ring of hoary tomentum. Fruit ripening about the middle of September and soon falling, on stout pedicels, in drooping narrow clusters, short-oblong, full and rounded at the ends, dark crimson, lustrous, marked by minute pale dots, usually about ⅝′ long and ½′ in diameter; calyx enlarged and persistent, the lobes elongated, strongly incurved, often deciduous before the fruit ripens; flesh thick, dry and mealy; nutlets 3, narrowed and acute at the ends, prominently ridged on the back with a high broad ridge, ⅓′ long.

A tree, rarely more than 15° high, with a straight trunk 5′—6′ in diameter, covered with thin bark separating into papery lustrous pale scales, stout branches forming a broad open irregular head, and slender glabrous branchlets bright chestnut-brown during their first season, becoming ashy gray the following year, and armed with many thick straight or curved bright chestnut-brown or red-brown spines 1′—1½′ long.

Distribution. Low hills and limestone ridges; western and southern Vermont; southern Connecticut (rocky shore of Alewive Creek, Waterford, New London County), and eastern and central New York (Whitesboro, Oneida County).

64. Cratægus lucorum Sarg.

Leaves broad-ovate to obovate or rarely oval, broad-cuneate or rounded at the entire base, coarsely serrate above with straight teeth tipped with large persistent bright red glands, and deeply divided above the middle into 3 or 4 pairs of wide acute or acuminate lobes, rather more than a third grown when the flowers open early in May and then light yellow-bronze color, covered on the upper surface with short soft pale hairs and glabrous on the lower surface, and at maturity membranaceous, smooth, dark dull green and glabrous above, pale yellow-green below, about 2′ long and 1¼′ wide, with a slender yellow midrib, and 3 or 4 pairs of thin primary veins extending obliquely to the point of the lobes; petioles slender, glandular, often somewhat winged toward the apex, 1′—1½′ in length; leaves at the end of vigorous shoots usually ovate and rounded at the broad base, more deeply lobed, and sometimes 3′ long and broad. Flowers ¾′ in diameter, on thin pedicels, in narrow compact few-flowered small villose corymbs; calyx broadly obconic, glabrous, the lobes narrow, acuminate, coarsely glandular-serrate, villose on the inner surface; stamens 20; anthers small, dark purple; styles 4 or 5. Fruit ripening about the middle of September and soon falling, on short stout pedicels, in erect few-fruited slightly villose clusters, obovoid until nearly fully grown and then short-oblong or somewhat obovoid, full and rounded at the ends, crimson, lustrous, marked by small pale dots, ½′—⅝′ long; calyx enlarged, the lobes elongated, coarsely glandular-serrate, villose above, closely appressed, often deciduous before the fruit ripens; flesh thick, yellow, dry and mealy; nutlets 4 or 5, thin, rounded, and sometimes obscurely ridged on the back, about ¼′ long.

A tree, 20°—25° high, with a tall straight trunk 6′—8′ in diameter, covered with close dark red-brown bark, slender ascending branches forming a narrow open head, and thin branchlets dark green and somewhat villose when they first appear, becoming dull orange-brown in their first summer and ultimately dark gray-brown, and armed with straight or slightly curved bright red-brown lustrous spines 1′—1½′ long.

Distribution. Rich moist soil along the margins of Oak-groves on the banks of sloughs; Barrington, Cook County, Illinois; near Ithaca, Thompkins County, New York.

65. Cratægus lacera Sarg.

Leaves rhombic to broad-ovate or rarely obovate, acute at apex, broadly cuneate and entire at base, coarsely often doubly serrate above with straight glandular teeth, and divided above the middle into numerous acute lobes, when they unfold coated below with thick hoary tomentum and villose above, nearly fully grown when the flowers open about the 20th of April and then glabrous on the lower surface and covered on the upper surface with short scattered pale hairs, and at maturity glabrous, light yellow-green, paler below than above, thin, about 1½′ long and 1¼′ wide, with a slender yellow midrib and few remote primary veins; petioles slender, villose, becoming glabrous or puberulous, slightly winged at the apex, often red toward the base, ¼′—⅓′ in length; leaves at the end of vigorous shoots broad-ovate, often deeply 3-lobed, coarsely serrate, 3′—4′ long and broad. Flowers ¾′ in diameter, on slender villose pedicels, in sparingly villose few-flowered corymbs; calyx-tube narrowly obconic, glabrous, the lobes linear-lanceolate, elongated, coarsely glandular-serrate, glabrous on the outer surface, villose on the inner surface; stamens 20; anthers small, rose color; styles 4 or 5. Fruit ripening toward the end of October, on short stout glabrous pedicels, in erect few-fruited clusters, ellipsoidal, rounded at the ends, bright cherry-red, lustrous, marked by occasional large dark dots, about ½′ long; calyx only slightly enlarged, with small nearly triangular villose spreading lobes mostly deciduous before the fruit ripens; flesh thick, orange color; nutlets 3—5, thin, narrowed at the ends, only slightly ridged on the rounded back, /16′ long.

A slender tree, 25°—30° high, with a tall trunk 4′—5′ in diameter, covered with pale scaly bark, small short branches forming a narrow head, and slender branchlets dark olive-green and villose when they first appear, becoming light red-brown and glabrous during their first summer, and ultimately dull light gray, and armed with thin straight bright chestnut-brown lustrous spines ¾′—1¾′ long.

Distribution. Low rich forest-glades near Fulton, Hempstead County, Arkansas.

66. Cratægus depilis Sarg.

Leaves ovate, acute or acuminate, rounded or broad-cuneate and often unsymmetrical at the entire base, sharply doubly serrate above with straight glandular teeth, and often divided into 4 or 5 pairs of short acute lobes, when they unfold deeply tinged with red and covered above with fine short caducous hairs, nearly half grown when the flowers open during the second week of May, and at maturity membranaceous, glabrous, smooth, yellowish to bluish green on the upper surface, pale on the lower surface, 1½′—2′ long, and 1′—1¼′ wide, with a slender midrib and 5 or 6 pairs of thin primary veins; turning yellowish and brown or russet color in the autumn; petioles slender, glabrous, sparingly glandular with minute glands, ¾′—1′ in length; leaves at the end of vigorous shoots often 2½′ long and 1½′ wide. Flowers ¾′ in diameter, on slender pedicels, in broad glabrous 8—12-flowered corymbs, with linear or oblong glandular bracts and bractlets; calyx narrowly obconic, glabrous, the lobes lanceolate, glandular-serrate, deeply tinged with purple; stamens 20; anthers pale rose color; styles 4 or 5. Fruit ripening early in September and soon falling, on slender pedicels, in drooping few-fruited clusters, subglobose to broad-obovoid, dark red to reddish purple, lustrous, ½′—¾′ long, and ⅜′—⅝′ in diameter; calyx only slightly enlarged, the lobes reflexed, glandular-serrate, and red on the upper side toward the base; flesh thick, yellow, sweet, juicy and slightly acid; nutlets 4 or 5, full and rounded at apex, narrowed and acute at base, and prominently but irregularly ridged on the back with a high sometimes grooved ridge, ¼′—/16′ long.

A tree, 20°—25° high, with a trunk 4′—8′ in diameter and 6°—9° long, covered with dark gray or gray-brown flaky bark, spreading branches forming an oblong or rounded open head, and slender glabrous branchlets bright red-brown and very lustrous during their first summer, becoming light gray-brown the following year, and armed with stout or slender nearly straight spines ¾′—1½′ long.

Distribution. Rich clay or gravelly soil in pastures and on the borders of woods; northeastern Illinois (Lake, Cook and Mill Counties).

67. Cratægus basilica Beadl.

Leaves ovate, acute or acuminate, broad-cuneate or rounded at the entire or crenate base, sharply and often doubly serrate above with straight slender glandular teeth, and divided into numerous short acute lateral lobes, more than half grown when the flowers open early in May and then roughened above by short pale hairs and glabrous below, and at maturity thin but firm in texture, bright green and scabrate above, paler below, 2½′—3′ long, and 1½′—2′ wide, with a slender yellow midrib, and thin veins arching to the point of the lobes; turning yellow and brown in the autumn; petioles slender, slightly winged at apex, 1′—1½′ in length. Flowers ½′—⅝′ in diameter, on elongated slender pedicels, in 5—15-flowered glabrous compact corymbs; calyx-tube broadly obconic, glabrous, the lobes slender, acuminate, glabrous, entire or occasionally serrate; stamens 15—20; anthers dark rose color; styles 3—5. Fruit ripening and falling early in September, on slender pedicels, in few-fruited drooping clusters, subglobose, scarlet, covered with a glaucous bloom, ½′—⅝′ in diameter; flesh soft, sweet, and edible; nutlets 3—5, narrowed and acute at the ends, prominently ridged on the back with a high broadly grooved ridge, ¼′—/16′ long.

A tree, sometimes 20° high, with a trunk 7′—8′ in diameter, covered with dark gray or brown scaly bark, ascending or slightly spreading branches forming a narrow irregular head, and stout glabrous branchlets dark chestnut-brown in their first season becoming dark gray, and armed with numerous slender bright chestnut-brown lustrous ultimately gray spines 2′—2½′ long.

Distribution. Open woods and the borders of fields and roads, western North Carolina, usually at altitudes of 2000°—3000° above the sea.

VIII. Molles.

Conspectus of the Arborescent Species.

Stamens 20.

Anthers pale yellow or white (rose color in 71).

Leaves broad and rounded, truncate or cordate at base; fruit subglobose to short-oblong or obovoid, red, crimson or scarlet.

Mature leaves glabrous on the upper surface.

Leaves thin.

Fruit subglobose to short-oblong, scarlet, ripening in August and September.

68. C. mollis (A).

Fruit obovoid to short-oblong, dark red, ripening in October.

69. C. sera (A).

Leaves subcoriaceous; fruit short-oblong to obovoid, crimson, ripening in October and November.

70. C. arkansana (C).

Mature leaves scabrate on the upper surface; fruit depressed-globose, red, ripening in August and September.

71. C. gravida (A).

Leaves broad-cuneate or rounded at base, acute or acuminate, scabrate on the upper surface at maturity.

Fruit red.

Leaves villose below at maturity on midrib and veins, those at the end of vigorous shoots cuneate at base; flowers in usually 7—12-flowered corymbs; fruit short-oblong, orange-red.

72. C. invisa (C).

Leaves hoary-tomentose below at maturity, those at the end of vigorous shoots rounded, cordate or abruptly cuneate at the broad base; flowers in 15—20-flowered corymbs; fruit ellipsoidal, ovoid, short-oblong or subglobose, crimson.

73. C. limaria (C).

Fruit bright canary yellow, subglobose; leaves villose below at maturity elliptic to ovate, oval or slightly obovate.

74. C. viburnifolia (C).

Leaves narrowed at base.

Mature leaves glabrous on the upper surface; fruit short-oblong to subglobose.

Leaves oblong-obovate or oval.

75. C. Berlandieri (C).

Leaves elliptic to ovate or slightly obovate.

76. C. meridionalis (C).

Mature leaves scabrate on the upper surface; fruit subglobose to short-oblong, red.

Leaves ovate to oval; flowers in 3—10-flowered corymbs; calyx-lobes glabrous.

77. C. Treleasei (C).

Leaves ovate; flowers in many-flowered corymbs; calyx-lobes villose.

78. C. canadensis.

Anthers rose color.

Leaves broad at base.

Mature leaves smooth on the upper surface.

Leaves thick, ovate, acute at apex; fruit short-oblong to obovoid, bright cherry red.

79. C. corusca (A).

Leaves thin, broad-ovate to suborbicular, rounded at apex; fruit subglobose to ovoid, bright yellow.

80. C. Kelloggii (A).

Mature leaves scabrate on the upper surface, oblong-obovate; fruit short-oblong, crimson.

81. C. induta (C).

Leaves narrowed at base; fruit red.

Leaves yellow-green.

Mature leaves glabrous on the upper surface; fruit short-oblong to obovoid.

82. C. texana (C).

Mature leaves scabrate on the upper surface.

Fruit subglobose to short-oblong.

83. C. quercina (C).

Fruit obovoid.

84. C. pyriformis (C).

Leaves blue-green, subcoriaceous, ovate to suborbicular, scabrate on the upper surface; fruit subglobose to short-oblong, red.

85. C. lanuginosa (C).

Stamens 10.

Anthers yellow.

Leaves broad at base.

Leaves smooth on the upper surface.

Leaves ovate or rarely oval, dark yellow-green above; fruit subglobose, crimson, ripening late in August.

86. C. arnoldiana (A).

Leaves ovate, blue-green above; fruit obovoid to short-oblong, scarlet, ripening in September.

87. C. champlainensis (A).

Leaves scabrate on the upper surface, ovate, acute, rounded or abruptly cuneate at base; anthers nearly white; fruit short-oblong, bright orange-red.

88. C. pennsylvanica (A).

Leaves cuneate at base, scabrate on the upper surface, ovate, acute; fruit obovoid, orange-red.

89. C. submollis (A).

Anthers rose color.

Leaves broad at the rounded, abruptly cuneate or cordate base.

Leaves scabrate on the upper surface.

Leaves oval, rounded or cuneate at base; flowers in wide many-flowered corymbs; fruit short-oblong, crimson.

90. C. Ellwangeriana (A).

Leaves oblong-ovate; flowers in compact few-flowered corymbs; fruit obovoid to short-oblong, scarlet.

91. C. Robesoniana (A).

Leaves smooth on the upper surface at maturity, ovate, usually broad-cuneate at base; fruit obovoid to short-oblong, crimson.

92. C. anomala (A).

Leaves cuneate at base, smooth on the upper surface at maturity; fruit subglobose, orange-red.

93. C. noelensis (C).

68. Cratægus mollis Scheele. Red Haw.

Leaves broad-ovate, acute, usually cordate or rounded at the wide base, coarsely and generally doubly serrate with straight glandular teeth, and more or less deeply divided into 4 or 5 pairs of acute or rounded lateral lobes, covered above with short pale hairs and hoary-tomentose below when they unfold, about half grown when the flowers open early in May and then membranaceous, light yellow-green and hairy above and pubescent or tomentose below, and at maturity firm in texture, dark yellow-green and slightly rugose on the upper surface and paler and pubescent or puberulous on the lower surface along the stout midrib, and 4 or 5 pairs of primary veins extending to the point of the lobes, 3′—4′ long and broad; petioles stout, terete, at first tomentose, ultimately pubescent or nearly glabrous, often slightly glandular with small dark caducous glands, 1′—1¼′ in length; leaves at the end of vigorous shoots more deeply lobed, with a deeper basal sinus, and frequently 5′—6′ long and broad. Flowers 1′ in diameter, on stout densely villose pedicels, in broad many-flowered tomentose corymbs, with conspicuous bracts and bractlets; calyx-tube narrowly obconic, hoary-tomentose, the lobes narrow, acuminate, coarsely glandular-serrate with bright red glands, villose on the outer, tomentose on the inner surface; stamens 20; anthers large, light yellow; styles 4 or 5, surrounded at base by a broad ring of hoary tomentum. Fruit ripening late in August and early in September, on stout pedicels, in drooping few-fruited villose clusters, short-oblong to subglobose, rounded at the ends, more or less pubescent, scarlet marked by occasional large dark dots; ¾′—1′ in diameter; calyx prominent, hairy, with large erect and incurved lobes usually deciduous before the fruit ripens; flesh thick, yellow, subacid, dry and mealy; nutlets 4 or 5, thin, rounded and obscurely ridged on the back, light brown, ¼′ long.

A tree, sometimes 40° high, with a tall trunk often 18′ in diameter, heavy wide-spreading smooth ashy gray branches forming a broad round-topped and often symmetrical head, and stout branchlets covered at first with a thick coat of long white matted hairs, villose during their first season, becoming glabrous in their second year, and armed with occasional straight thick bright chestnut-brown shining spines 1′—2′ long.

Distribution. Low rich soil usually on the bottom-lands of streams; northern Ohio and southwestern Ontario (Point Edward) to northern Missouri, eastern South Dakota, eastern Nebraska, and eastern Kansas; common; near Nashville, Davidson County, Tennessee.

69. Cratægus sera Sarg.

Leaves oblong-ovate, acute or acuminate, rounded, truncate or slightly cordate at the broad base, irregularly divided into 4 or 5 pairs of short acute lateral lobes, and sharply and sometimes doubly serrate nearly to the base with straight glandular teeth, unfolding about the 1st of May with the opening of the flowers and then covered above with short soft white hairs and tomentose below, and at maturity membranaceous, dark yellow-green and glabrous on the upper surface, pubescent on the lower surface, 2′—4′ long, and 2½′—3′ wide, with a slender midrib, and thin remote primary veins extending to the point of the lobes; petioles slender, tomentose, becoming pubescent, 1′—1½′ in length; leaves at the end of vigorous shoots more deeply lobed, and often 4′—5′ long and 3′—4′ wide. Flowers ¾′ in diameter, on stout densely villose pedicels, in compact many-flowered tomentose corymbs; calyx-tube broadly obconic, coated with broad matted pale hairs, the lobes broad, acute or acuminate, glandular-serrate with large dark glands, tomentose on the outer surface and villose on the inner surface; stamens 20; anthers pale yellow; styles 4 or 5, usually 5. Fruit ripening about the 1st of October, on stout puberulous or villose pedicels, in drooping or erect few-fruited clusters, obovoid or short-oblong, dull dark red, marked by small pale dots, usually slightly villose or pubescent at the ends, ⅔′ long, and ½′ in diameter; calyx enlarged, with erect, coarsely glandular-serrate, incurved lobes often deciduous before the ripening of the fruit; flesh thick, dry and mealy; nutlets usually 5, thin, light brown, irregularly grooved on the back with a broad shallow groove, ¼′ long.

A tree, 30°—40° high, with a tall straight trunk 12′—18′ in diameter, thick branches forming a broad round-topped symmetrical head, and branchlets hoary-tomentose at first, becoming light red-brown and puberulous and ultimately pale orange-brown, and armed with occasional straight or slightly curved chestnut-brown lustrous spines 1¼′—1½′ in length.

Distribution. Walpole Island, Lamberton County, southwestern Ontario; Belle Isle in the Detroit River, near Port Huron, St. Clair County, and in the neighborhood of Grand Rapids, Kent County, Michigan; northeastern Illinois (Cook, Will, Lake and Dupage Counties), and in the neighborhood of Milwaukee, Milwaukee County, Wisconsin.

70. Cratægus arkansana Sarg.

Leaves oblong-ovate or oval, acute, rounded, broadly cuneate or truncate at base, usually divided above the middle into 3 or 4 pairs of short broad acute lobes, and serrate sometimes to the base with short straight glandular teeth, when the flowers open about the middle of May nearly one third grown and coated with soft white hairs, and at maturity thick and leathery, dull dark green and glabrous on the upper surface, pale yellow-green on the lower surface, 2′—3′ long, and 1¾′—2′ wide, with a stout light yellow midrib and primary veins slightly villose below, conspicuous secondary veins and reticulate veinlets; late in October and in November turning bright clear yellow; petioles stout, deeply grooved, more or less winged toward the apex, glandular with minute usually deciduous dark glands, at first tomentose, ultimately glabrous or puberulous, turning dark red after midsummer, 1′—1½′ in length; leaves at the end of vigorous shoots broad-ovate, rounded or truncate at base, often 4′ long and 3′ wide. Flowers nearly 1′ in diameter, on short stout pedicels, in broad rather compact many-flowered villose corymbs; calyx-tube narrowly obconic, coated with long matted pale hairs, the lobes short, acute, coarsely glandular-serrate, glabrous or slightly villose; stamens 20; anthers large, pale yellow; styles 5. Fruit ripening at the end of October and falling gradually at the end of several weeks, on stout villose pedicels, in few-fruited drooping clusters, short-oblong or rarely obovoid, rounded and slightly tomentose at the ends, bright crimson, very lustrous, marked by few large dark dots, ¾′—1′ long, about ¾′ in diameter; calyx little enlarged, with small linear-lanceolate coarsely glandular-serrate erect and persistent lobes; flesh thick, yellow, subacid; nutlets 5, small in comparison to the size of the fruit, thin, rounded or slightly and irregularly ridged on the back, ⅓′ long.

A tree, 20° high, with a tall straight stem, thick slightly ascending wide-spreading branches forming a broad open irregular head, and stout branchlets dark green and covered early in the season with long pale hairs, becoming orange-brown, glabrous, and very lustrous in their first winter, and unarmed or armed with occasional straight light chestnut-brown shining spines, ⅓′—½′ in length.

Distribution. Bottom-lands of the White River near Newport, Jackson County, Arkansas; hardy as far north as eastern Massachusetts, and unsurpassed late in the autumn in the beauty of its large brilliant abundant fruits long persistent on the branches.

71. Cratægus gravida Beadl.

Leaves broad-ovate, acute, rounded or truncate at base, coarsely and often doubly serrate with incurved glandular teeth, and slightly incisely lobed, roughened above by short pale hairs and hoary-tomentose below when they unfold, nearly half grown when the flowers open about the 1st of May, and at maturity thin, firm, dark green, lustrous and scabrate above, paler and pubescent or puberulous below, particularly on the slender midrib and veins, 1¾′—2½′ long, and 1½′ wide; turning in the autumn yellow, orange and brown; petioles slender, tomentose early in the season, becoming pubescent or nearly glabrous, about ½′—1′ in length; leaves at the end of vigorous shoots oblong-ovate to nearly orbicular, round or cuneate at the broad base, more coarsely serrate, more deeply lobed, and often 2½′—3′ long and wide, their petioles ¾′—1′ long. Flowers about ⅝′ in diameter, on short hoary-tomentose pedicels, in narrow crowded many-flowered hoary-tomentose corymbs; calyx-tube broadly obconic, covered with matted pale hairs, the lobes gradually narrowed, acuminate, glandular-serrate, villose; stamens 20; anthers dark rose; styles 5. Fruit ripening in August and September, on elongated tomentose pedicels, in few-fruited drooping clusters, depressed-globose, red; calyx enlarged, the lobes conspicuously serrate, puberulous on the upper surface, reflexed and closely appressed, sometimes deciduous from the ripe fruit; flesh thin, yellow, dry and mealy; nutlets 5, thin, narrow and rounded at base, acute at apex, rounded and obscurely grooved on the back, about /16′ long.

A tree, sometimes 20° high, with a trunk 8′—10′ in diameter, heavy wide-spreading branches forming a broad round-topped head, and stout branchlets covered at first with a thick coat of matted pale hairs, orange-red and puberulous at the end of their first season, glabrous and reddish brown the following year, and armed with slender nearly straight spines about 1½′ long.

Distribution. Limestone hills in the neighborhood of Nashville, Davidson County, Tennessee.

72. Cratægus invisa Sarg.

Leaves ovate to oval, acute or acuminate at apex, cuneate or rounded at base, coarsely often doubly serrate with broad straight glandular teeth, and slightly divided usually only above the middle into 3 or 4 pairs of small acuminate lobes, densely tomentose below and villose above when they unfold, about one third grown when the flowers open at the end of March and then thin, dark yellow-green and roughened on the upper surface by short hairs and coated below with long matted white hairs, and at maturity thin, yellow-green, scabrate and lustrous above, hairy below especially on the midrib and veins, 2½′—3′ long, and 2′—2½′ wide; petioles slender, slightly wing-margined at apex, covered with pale hairs early in the season, becoming nearly glabrous, and 1½′—2′ in length; leaves at the end of vigorous shoots broad-ovate, acuminate, abruptly cuneate at the wide base, more coarsely serrate, deeply divided into acute lateral lobes, and often 3½′—4′ long and 3′—3½′ wide; petioles slender, villose, 1½′—2′ in length. Flowers opening at the end of March, about ¾′ in diameter, on slender pedicels thickly coated like the wide calyx-tube with long matted white hairs, in broad mostly 7—12-flowered corymbs; calyx-lobes gradually narrowed from the base, short, broad, acuminate, laciniately glandular-serrate, thickly covered with long white hairs on the outer surface, villose above the middle on the inner surface; stamens 20; anthers white; styles 3—5, surrounded at base by a ring of long white hairs. Fruit ripening at the end of October, on long slender slightly hairy pedicels, in erect or spreading few-fruited clusters, short-oblong, full and rounded and slightly hairy at the ends, orange-red, marked by large pale dots, and about ½′ in diameter; calyx little enlarged, with spreading lobes dark red on the upper side below the middle and villose toward the apex; flesh thin, yellow, dry and mealy; nutlets 3—5, rounded at the ends, broader at apex than at base, rounded and only slightly grooved on the back, ⅕′ long.

A tree, sometimes 30° high, with a tall trunk covered with dark brown bark broken into small closely appressed plate-like scales, large spreading branches forming a wide irregular head, and stout slightly zigzag branchlets clothed when they first appear with hoary tomentum, dull gray-brown, marked by small pale lenticels and slightly pubescent at the end of their first season and dark gray the following year, and unarmed or armed with occasional slender straight chestnut-brown spines 1′—1¼′ long.

Distribution. In dense woods on the rich bottom-lands of Red River near Fulton, Hempstead County, and near Texarkana, Miller County, Arkansas.

73. Cratægus limaria Sarg.

Cratægus Mackensenii Sarg.

Leaves ovate, acute, concave-cuneate or rounded at base, coarsely often doubly serrate with broad straight glandular teeth, and slightly divided into 3 or 4 pairs of small acute lateral lobes, not more than a quarter grown when the flowers open early in April and then thin, yellow-green and covered above with short white hairs and thickly coated below with hoary tomentum, and at maturity light green and scabrate on the upper surface, pale and tomentose on the lower surface, 2½′—3′ long, and 1½′—2′ wide, with a stout midrib and thin primary veins; petioles slender, slightly wing-margined at apex, covered when they first appear with long matted white hairs, villose through the season, and 1′—1½′ in length; leaves at the end of vigorous shoots broad-ovate, rounded or cordate at the wide base, more deeply lobed, and often 4′ long and broad. Flowers opening early in April, ⅘′—1′ in diameter, on long slender pedicels coated with matted white hairs, in compact 15—20-flowered villose corymbs; calyx-tube broadly obconic, thickly covered with white hairs, the lobes gradually narrowed from the base, wide, acuminate, laciniately glandular-serrate, villose; stamens 20; anthers white; styles 3—5, surrounded at base by a narrow ring of pale tomentum. Fruit ripening in October, on long stout erect or spreading hairy pedicels, in few-fruited clusters, ellipsoidal to ovoid or short-oblong, rounded at apex, truncate at base, crimson, lustrous, marked by large pale dots, villose especially at the ends, ½′—⅗′ in diameter; calyx prominent, with a long villose tube, and erect villose persistent lobes dark red on the upper side below the middle, their tips slightly spreading or incurved; flesh thick, yellow, dry and mealy; nutlets 3—5, narrowed and rounded at apex, rounded at the broad base, slightly grooved on the back, ⅕′—¼′ long.

A tree, often 30° high, with a tall trunk 8′—12′ in diameter, covered with dark scaly bark, stout ascending branches forming a narrow irregular head, and slender zigzag branchlets thickly coated when they first appear with long-white hairs, light orange-brown, lustrous, pubescent and marked by pale lenticels at the end of their first season, dull gray-brown and glabrous the following year, and armed with slender straight or slightly curved purple ultimately ashy gray spines 2′—2½′ long.

Distribution. In dense woods on the rich bottom-lands of the Red River near Fulton, Hempstead County, Arkansas; river banks; western Texas (Guadalupe River, near Victoria, Victoria County; Cibolo River, Sutherland Springs, Wilson County; San Antonio River, Bexar County; C. Mackensenii Sarg.).

74. Cratægus viburnifolia Sarg.

Leaves elliptic to ovate, oval or slightly obovate, acute or rounded at apex, concave-cuneate at the entire base, coarsely often doubly serrate above with straight glandular teeth, and slightly and irregularly divided above the middle into 2 or 3 pairs of small acute lobes, half grown when the flowers open about the 20th of March and then thin, yellow-green and roughened above by short white hairs and hoary-tomentose below, and at maturity thick, deep green, very lustrous and scabrate on the upper surface, coated on the lower surface with pale hairs, 2½′—3½′ long, and 2′—2½′ wide, with a prominent midrib and primary veins; petioles slightly wing-margined at apex, densely hoary-tomentose early in the season, becoming glabrous, ⅖′—1½′ in length. Flowers about ¾′ in diameter, on long slender tomentose pedicels, in wide lax mostly 5—12-flowered corymbs, with large lanceolate to spatulate foliaceous bracts and bractlets slightly serrate above the middle, and generally persistent until after the petals fall; calyx-tube narrowly obconic, thickly coated with matted white hairs, the lobes gradually narrowed from the base, long, slender, acuminate, laciniately glandular-serrate, slightly villose on the outer surface, densely villose on the inner surface; stamens 20; anthers white; styles 4 or 5. Fruit ripening early in October, on long slender drooping slightly hairy pedicels, in few-fruited clusters, subglobose, bright canary yellow, about 1′ in diameter; calyx little enlarged, with spreading lobes; flesh thick, light yellow, soft and succulent; nutlets 4 or 5, gradually narrowed and rounded at the ends, irregularly ridged on the back with a broad grooved ridge, ⅓′ long.

A tree, 30°—35° high, with a tall trunk sometimes a foot in diameter, covered with gray scaly bark, large ascending and spreading branches forming an open irregular head, and stout nearly straight unarmed branchlets thickly coated with hoary tomentum when they first appear, becoming purple, lustrous and nearly glabrous at the end of their first season and dark brown or gray-brown the following year.

Distribution. Borders of woods in low ground, valley of the Brazos River near Columbia, Brazoria County, and in low woods on the Colorado River, at Wharton, Wharton County, Texas.

75. Cratægus Berlandieri Sarg.

Leaves oblong-obovate or oval, acute or acuminate, gradually narrowed, cuneate and entire below the middle, coarsely and often doubly serrate with broad straight or incurved glandular teeth, and unequally divided above into numerous acute or acuminate lobes, when the flowers open from the middle to the end of March coated on the upper surface with short pale caducous hairs and on the lower surface with thick hoary tomentum, and at maturity thin and firm in texture, glabrous, dark green, and lustrous above, pale and pubescent below, and usually about 3′ long and 2′ wide, with a slender midrib, remote primary veins extending to the point of the lobes, conspicuous secondary veins, and reticulate veinlets; petioles more or less winged toward the apex, tomentose early in the season, becoming pubescent, ½′—¾′ in length; leaves at the end of vigorous shoots often 5′ long and 3′ wide, with rounded, acute lobes. Flowers ¾′ in diameter, on long stout hoary-tomentose pedicels, in broad loose many-flowered tomentose corymbs, with oblong-obovate to lanceolate finely glandular-serrate villose conspicuous bracts and bractlets; calyx-tube broadly obconic, covered with thick pale tomentum, the lobes broad, acute, very coarsely glandular-serrate, tomentose on the outer surface and villose on the inner surface; stamens 20, anthers yellow; styles 5, surrounded at base by tufts of white hairs. Fruit ripening after the middle of October, on slender elongated pedicels, in loose dropping clusters, short-oblong to subglobose, scarlet, about ½′ long; calyx much enlarged, with coarsely serrate erect and persistent villose lobes; flesh thin, yellow, dry and mealy; nutlets 5, rounded and occasionally obscurely grooved on the back, about ¼′ long.

A tree, 15°—20° high, with a tall straight trunk 8′—10′ in diameter, covered with thin dark brown furrowed bark, spreading branches forming a broad open head, and branchlets hoary-tomentose at first, soon puberulous, dull reddish brown or yellow-brown by midsummer, becoming ashy gray late in the autumn, and armed with few straight gray spines about 1′ in length.

Distribution. Low rich woods on the bottom-lands of the Brazos River at Columbia and Brazoria, Brazoria County, Texas.

76. Cratægus meridionalis Sarg.

Leaves elliptic to ovate or slightly obovate, acuminate, cuneate at the entire base, and coarsely often doubly serrate above with broad straight glandular teeth, coated below with hoary tomentum and covered above with short white hairs when they unfold, more than half grown when the flowers open from the first to the middle of April, and at maturity thin, yellow-green and scabrate on the upper surface, paler and villose-pubescent on the lower surface, especially on the slender midrib and primary veins, 2′—3½′ long, and 1′—2′ wide; petioles slender, slightly wing-margined at apex, densely villose-pubescent with white hairs early in the season, becoming glabrous or nearly glabrous, ½′—¾′ in length; leaves at the end of vigorous shoots broad-ovate to broad-elliptic, more coarsely serrate, occasionally slightly divided into short broad lateral lobes, often 4′ long and 2½′ wide, with a stout midrib and petioles broadly wing-margined at apex, and about ½′ in length. Flowers ¾′ in diameter, on stout pedicels thickly covered like the narrow obconic calyx-tube with matted silvery white hairs, in broad compact many-flowered villose corymbs, with conspicuous glandular-serrate villose bracts and bractlets mostly persistent until after the flowers open; calyx-lobes narrow, acuminate, laciniately glandular-serrate, slightly villose-pubescent when the buds open; stamens 20; anthers white; styles 3—5, surrounded at base by a broad ring of white tomentum. Fruit ripening from the middle to the end of September, on elongated slender puberulous pedicels, in few-fruited drooping red-stemmed clusters, short-oblong to subglobose, rounded at the ends, scarlet, ½′ to ⅔′ in diameter, the calyx persistent, much enlarged, with erect or spreading conspicuous lobes; nutlets 3—5, rounded at base, acute at apex, ridged on the back with a high rounded ridge, about ⅓′ long.

A tree, often 25° high, with a trunk 8′ in diameter, covered with dark bark slightly divided by shallow fissures into broad thin plates, spreading ashy gray branches forming a round-topped head, and slender zigzag branchlets, covered when they first appear with long white hairs, soon glabrous, orange-brown or reddish brown during their first season and dull gray the following year, and armed with numerous straight slender purple spines 1′—2′ in length.

Distribution. Limestone soil, in upland woods and glades; common in the limestone belt of central Alabama, from the neighborhood of Gallion, Hale County to western Mississippi (Starkville, Oktibbeha County, and Brookville, Noxubee County).

77. Cratægus Treleasei Sarg.

Leaves ovate to elliptic, acute, concave-cuneate or rounded at the narrow base, sharply doubly serrate above with straight glandular teeth, and slightly divided into 3 or 4 pairs of narrow acuminate lateral lobes, unfolding with the opening of the flowers at the end of April or early in May and then light yellow-green tinged with bronze color, lustrous and covered above with short shining caducous white hairs and hoary-tomentose below, and at maturity thin, light yellow-green and scabrate on the upper surface, paler and pubescent on the lower surface, especially on the slender midrib, and 4 or 5 pairs of thin primary veins extending obliquely to the point of the lobes, 1¾′—2¼′ long, and 1½′—2′ wide; petioles slender, more or less wing-margined at apex, villose early in the season, pubescent in the autumn, ½′—¾′ in length; leaves at the end of vigorous shoots broad-ovate, acute, cuneate at the wide base, often 2½′—3′ long and 2′—2½′ wide; petioles stout, wing-margined at apex ¾′—1′ long. Flowers 1′ in diameter, on short stout pedicels covered with matted pale hairs, in 3—10-flowered compact compound or rarely simple villose corymbs; calyx-tube broadly obconic, covered with matted pale hairs, the lobes glabrous, narrowed from the base, with wide rounded sinuses between them, slender, acuminate, tipped with a small red gland, and glandular-serrate with stipitate red glands; stamens 20; anthers pale yellow; styles 4 or 5, usually 5. Fruit ripening at the end of September, on stout erect villose pedicels, in few-fruited clusters, subglobose, often broader than high, crimson, lustrous, marked by numerous large pale dots, pubescent at the ends, and ½′—¾′ in diameter; calyx prominent, with a short villose tube, and reflexed appressed villose lobes often deciduous from the ripe fruit; flesh thick, light yellow, dry and mealy; nutlets 4 or 5, thin, full and rounded at apex, narrowed and acute at base, grooved with a broad shallow groove and sometimes irregularly ridged on the back, about /16′ long.

A tree, 20°—25° high, with a tall trunk sometimes 6′ in diameter, slender branches forming a narrow open head, and thin nearly straight branchlets thickly covered at first with long lustrous white hairs, dull light reddish brown and puberulous at the end of their first season, becoming dark gray-brown, and armed with stout straight or slightly curved dark purple shining spines usually about 1¼′ long, or unarmed.

Distribution. Banks of small streams in moist soil from Doe Run to Bismarck, St. François County, Missouri.

78. Cratægus canadensis Sarg.

Leaves ovate, short-pointed, slightly lobed usually only above the middle with short broad acute lobes, and coarsely and frequently doubly serrate to the broad-cuneate base with spreading glandular teeth, coated above in early spring with soft white hairs, and below with dense hoary tomentum, about a third grown when the flowers open at the end of May, and at maturity thin and firm in texture, blue-green and scabrate on the upper surface, pale and pubescent on the lower surface on the midrib and primary veins, 2′—2½′ long, and 1½′ to nearly 3′ wide; petioles slender, glandular, often more or less winged above, at first tomentose, becoming nearly glabrous, ¾′—1′ in length; leaves at the end of vigorous shoots broad-ovate, truncate or slightly cordate at the broad base, more deeply lobed, often 2½′—3′ long and wide, the petioles wing-margined at apex often glandular, and 1′—1½′ in length. Flowers about ¾′ in diameter, in broad loose tomentose corymbs; calyx-tube broadly obconic, villose with long matted hairs, the lobes lanceolate, villose, and glandular with large red stipitate glands; stamens 20; anthers small, nearly white; styles 5, surrounded at base by a thin ring of pale tomentum. Fruit ripening early in October and falling gradually until after midwinter, on stout pedicels, in erect slightly villose few-fruited clusters, short-oblong to subglobose, crimson, lustrous, marked by large scattered pale dots, slightly hairy toward the ends, ½′—⅝′ long, ⅓′—½′ in diameter; calyx prominent, the lobes gradually narrowed from a broad base, elongated, glandular, villose, spreading or reflexed, often deciduous before the fruit ripens; flesh thin, pale yellow, dry and mealy; nutlets 5, thin, rounded and irregularly ridged on the back, ¼′ long.

A tree, 18°—30° high, with a trunk 6′—8′ in diameter, stout spreading branches forming a broad round-topped symmetrical head, and stout zigzag branchlets dark green and covered with matted pale hairs when they first appear, soon becoming light orange-brown and very lustrous, and armed with numerous stout straight or slightly curved dark chestnut-brown shining spines 2′—2½′ long.

Distribution. Limestone ridges near the St. Lawrence River at Châteaugay, Caughnawaga, and La Tortue in the Province of Quebec.

79. Cratægus corusca Sarg.

Leaves ovate, acute, truncate, rounded or slightly cordate at the broad base, regularly divided into 4 or 5 pairs of short acute lateral lobes, and doubly serrate with straight glandular teeth, when they unfold covered above with short soft pale hairs and glabrous below, about a third grown when the flowers open the middle of May, and at maturity thin but firm and rigid in texture, glabrous, dark yellow-green, bright and lustrous above, pale yellow-green below, 2′—2½′ long and wide, with a slender pale midrib and primary veins; petioles slender, villose early in the season, soon becoming glabrous and dark red below the middle, 1½′—2½′ in length; leaves at the end of vigorous shoots frequently divided into narrow acute lateral lobes, and often 3½′—4′ long and wide. Flowers ¾′ in diameter, on stout villose pedicels, in compact narrow many-flowered corymbs covered with matted pale hairs; calyx-tube narrowly obconic, glabrous, or villose toward the base, the lobes narrowed from a broad base, acute, coarsely glandular-serrate, villose on the inner surface; stamens 20; anthers small, pale pink; styles 4 or 5. Fruit beginning to ripen and fall about the middle of September and continuing to fall until the end of October, on stout pedicels, in glabrous few-fruited clusters, short-oblong to obovoid, bright cherry-red, lustrous, marked by dark scattered pale dots, ⅝′—¾′ long, and ½′—⅝′ in diameter; calyx little enlarged, the lobes slightly glandular-serrate, usually deciduous before the fruit ripens; flesh thick, yellow, dry and mealy; nutlets 4 or 5, dark-colored, rounded on the back, ¼′ long.

A tree, 18°—20° high, with a tall trunk 8′—10′ in diameter, wide-spreading branches forming a handsome symmetrical head, and stout branchlets dark green and coated with matted pale hairs when they first appear, soon becoming light red-brown, light orange-brown and lustrous in their second year, and armed with thick nearly straight bright chestnut-brown spines often 3′ in length.

Distribution. Sandy shores of Lake Zurich, Lake County, Illinois.

80. Cratægus Kelloggii Sarg.

Leaves broad-ovate to suborbicular, rounded and often short-pointed at apex, rounded, broadly cuneate or truncate at the entire base, coarsely serrate above with straight gland-tipped teeth, and divided usually only above the middle into several short broad acute or acuminate lobes, about half grown when the flowers open during the last week of April and then thin, yellow-green, covered above with short pale hairs and pubescent below on the midrib and veins, and at maturity thin but firm in texture, dark yellow-green, glabrous and smooth on the upper surface, pale and glabrous on the lower surface with the exception of a few hairs near the base of the thin yellow midrib and of the 4 or 5 pairs of slender prominent primary veins arching to the point of the lobes, 2′—2½′ long, 1¾′—2½′ wide, and often broader than long; petioles slender, slightly winged at apex, villose while young with long matted white hairs, becoming glabrous, ¾′—1′ in length. Flowers ⅝′ in diameter, on slender hairy pedicels, in compact 5—10-flowered villose corymbs, with oblong-obovate to linear acuminate glandular bracts and bractlets mostly persistent until the flowers open; calyx-tube broadly obconic, slightly hairy at base, glabrous above, the lobes slender, acuminate, glandular with minute dark red stipitate glands, or entire, glabrous on the outer surface, sparingly villose on the inner surface; stamens 20; anthers pale rose color; styles 5. Fruit ripening at the end of September and soon falling, on long slender glabrous pedicels, in few-fruited drooping clusters, subglobose to short-ovoid, bright yellow, marked by many small pale dots, ¾′—1′ in diameter; calyx small, with spreading reflexed lobes slightly villose toward the apex and often deciduous from the ripe fruit; flesh thin, yellow, dry and mealy; nutlets 5, rounded and very slightly grooved on the back, about ⅜′ long.

A tree, 20°—25° high, with a tall trunk 4′—5′ in diameter, covered with nearly black deeply furrowed bark, erect branches, and nearly straight branchlets dark green tinged with red and slightly villose when they first appear, bright red-brown and lustrous at the end of their first season, becoming dark dull reddish brown the following year, and unarmed, or armed with slender nearly straight bright chestnut-brown shining spines usually about 1′ long.

Distribution. Banks of the Desperes River, South St. Louis, St. Louis County, Missouri; not common.

81. Cratægus induta Sarg. Turkey Apple.

Leaves oblong-obovate, acute, cuneate, rounded or rarely truncate at the broad entire base, coarsely doubly serrate above with glandular teeth, and slightly and irregularly divided into broad acute lateral lobes, about a third grown when the flowers open from the middle to the end of April and then thin, light yellow-green and roughened above by short lustrous white hairs and hoary-tomentose below, and at maturity thin, dark yellow-green and scabrate on the upper surface, pale and tomentose or pubescent on the lower surface, particularly on the stout midrib and 4 or 5 pairs of prominent primary veins, 3′—4′ long, and 2½′—3′ wide; petioles slender, more or less wing-margined at the apex, glandular, hoary-tomentose early in the season, becoming sparingly villose in the autumn, 1¼′—1½′ in length. Flowers ¾′ in diameter, on slender tomentose pedicels, in broad many-flowered hoary-tomentose corymbs; calyx-tube narrowly obconic, thickly coated with long densely matted white hairs, the lobes small, acuminate, glandular-serrate, villose; stamens 20; anthers small, rose color; styles 5, surrounded at base by a broad ring of snow-white hairs. Fruit ripening the middle of October, on stout villose pedicels, in few-fruited clusters, short-oblong, rounded and villose at the ends, crimson or reddish yellow, lustrous, marked by small pale dots, ¾′—2′ in diameter; calyx prominent, with a short tomentose tube and much enlarged coarsely glandular-serrate hairy erect incurved lobes often deciduous from the ripe fruit; flesh thick, orange-colored, with an astringent subacid flavor; nutlets 5, thin, rounded and slightly grooved on the back, /16′—⅜′ long.

A tree, sometimes 25° high, with a trunk often a foot in diameter, covered with thick dark brown furrowed bark, large spreading and ascending branches forming an open irregular head, and stout branchlets covered at first with long matted white hairs, light orange-brown, lustrous and puberulous at the end of their first season, becoming ashy gray or light grayish brown the following year, and armed with many stout nearly straight dark purple shining spines usually about 2½′ long.

Distribution. Dry upland woods, near Fulton, Hempstead County, Arkansas; common.

82. Cratægus texana Buckl.

Leaves broad-ovate, acute or rarely rounded at apex, broadly concave-cuneate at base, coarsely doubly glandular-serrate above, and usually divided above the middle into 4 or 5 pairs of broad acute lobes, covered above when they unfold with short soft pale hairs and below with a thick coat of hoary tomentum, more than half grown when the flowers open late in March, and at maturity thick and firm, dark green and lustrous above, pale and pubescent or tomentose below, particularly on the stout midrib, primary veins, prominent secondary veins and reticulate veinlets, 3′—4′ long, 2½′—3′ wide; petioles stout, deeply grooved, more or less winged above, at first tomentose, becoming nearly glabrous, ½′—¾′ in length; leaves at the end of vigorous shoots sometimes truncate or slightly cordate at the broad base, more deeply lobed, and frequently 3′ long and wide. Flowers ¾′ in diameter, on elongated slender densely villose pedicels, in broad open many-flowered tomentose corymbs, with oblong or oblong-obovate acute conspicuous villose bracts and bractlets often 1½′ in length; calyx-tube broadly obconic, coated with pale tomentum, the lobes foliaceous, gradually narrowed from a broad base, acuminate, coarsely glandular-serrate, and villose with long matted pale hairs; stamens 20; anthers large, dark red; styles 5, surrounded at base by a narrow ring of pale tomentum. Fruit ripening toward the end of October, in drooping many-fruited tomentose ultimately glabrous clusters, obovoid and tomentose until nearly grown, becoming when fully ripe short-oblong or slightly obovoid, rounded at the ends, bright scarlet, marked by occasional large pale dots, puberulous at apex, ¾′—1′ long; calyx enlarged, with glandular-serrate usually erect lobes, dark red at base on the upper side, often deciduous before the ripening of the fruit; flesh thick, yellow, sweet, and edible; nutlets 5, slightly grooved on the back, ¼′—⅓′ long.

A tree, often 30° high, with a tall trunk sometimes a foot in diameter, thick branches ascending while the tree is young, forming an open irregular crown, and spreading in old age into a broad symmetrical round-topped head, and branchlets dark bronze-green and covered with long matted white hairs when they first appear, becoming dull reddish brown and ultimately pale ashy gray, and armed with occasional thin nearly straight bright chestnut-brown lustrous spines usually about 2′ long, or often unarmed.

Distribution. Rich bottom-lands, Texas coast region; valley of the lower Brazos River to those of the Navidad (Canardo, Jackson County), Guadalupe (Victoria, Victoria County), and Cibolo (Sutherland Springs, Wilson County).

83. Cratægus quercina Ashe.

Leaves elliptic to obovate, usually acute or occasionally rounded at apex, obtusely or acutely cuneate at the entire base, irregularly doubly serrate above with slender glandular teeth, and often divided above the midrib into narrow acuminate lobes, when they unfold conspicuously plicate, often dark red and coated above with long soft pale hairs and covered below with a thick coat of silvery white shining tomentum, about a third grown when the flowers open from the middle to the end of March, and at maturity thin but firm in texture, dark green, lustrous and scabrate above, pale and pubescent or tomentose below, and 2′—2½′ long and wide, with a slender midrib, 4 or 5 pairs of thin primary veins, and conspicuous reticulate veinlets; petioles stout, tomentose, about ½′ in length; leaves at the end of vigorous shoots broad-ovate, rounded or obtusely cuneate at the wide base, usually deeply divided into numerous acuminate lateral lobes, often 3′ long and 2½′ wide. Flowers ¾′ in diameter, on long slender tomentose pedicels, in broad many-flowered lax hoary-tomentose corymbs, with oblong-obovate glandular-serrate villose bracts and bractlets; calyx-tube narrowly obconic, hoary-tomentose, the lobes short, acute, coarsely glandular-serrate, tomentose; stamens 20; anthers small, dark red; styles 5, surrounded at base by tufts of long snow-white hairs. Fruit ripening after the middle of October, on slender nearly glabrous pedicels, in few-fruited tomentose spreading clusters, subglobose but often rather longer than broad, rounded at the ends, tomentose until nearly fully grown, glabrous at maturity, dark red, marked by numerous large pale dots, about ½′ in diameter; calyx prominent, with short spreading often deciduous lobes; flesh thin, light yellow, hard and dry, generally shrivelling before the fruit falls; nutlets 5, rounded and ridged on the back, about ¼′ long.

A tree, remarkable for the lustre of its white tomentum, occasionally 25° high, with a tall trunk 6′—8′ in diameter, covered with light gray scaly bark, becoming near the base of old trees deeply furrowed and nearly black, ascending branches forming a broad symmetrical head, and branchlets coated when they first appear with hoary tomentum, becoming light red-brown and more or less villose during their first season, glabrous and rather darker in their second year, and armed with numerous straight or slightly curved chestnut-brown shining spines usually 1′—1¼′ long.

Distribution. Sandy bottom-lands in open Live Oak-forests on the Brazos River, near Columbia, Brazoria County, Texas.

84. Cratægus pyriformis Britt.

Leaves oval to broad-ovate, acute and often short-pointed at apex, gradually narrowed and concave-cuneate at the entire base, sharply and sometimes doubly serrate above with straight glandular teeth, and often slightly and irregularly lobed above the middle, fully grown when the flowers open about the 10th of May and then thin, light yellow-green, roughened above by short rigid pale hairs and pubescent below, particularly on the slender midrib and 5 or 6 pairs of remote primary veins, and at maturity thin and firm, lustrous and scabrate on the upper surface, pale and pubescent on the lower surface, and generally about 3′ long and 2′ wide; petioles slender, winged at apex, tomentose, ultimately pubescent, 1′—1¼′ in length; leaves at the end of vigorous shoots usually ovate, coarsely serrate, more deeply lobed, and frequently 4′—5′ long and 3′—4′ wide. Flowers 1′ in diameter, on long slender tomentose pedicels, in broad many-flowered lax corymbs; calyx-tube narrowly obconic, villose, the lobes narrow, acuminate, glandular-serrate, and covered more or less thickly with pale hairs; stamens 20; anthers pale rose color; styles 4 or 5, usually 5, surrounded at base by a broad ring of pale tomentum. Fruit ripening in October, on long slender pubescent pedicels, in drooping few-fruited clusters, obovoid, rounded at the ends, bright cherry-red, lustrous, marked by occasional large pale dots, about ⅝′ long and ½′ in diameter; calyx prominent, with linear glandular-serrate closely appressed lobes often deciduous before the fruit ripens; flesh thin, light yellow, juicy; nutlets 4 or usually 5, rounded, and deeply grooved on the back, dark brown, ⅝′ long.

A tree, 25°—30° high, with a trunk a foot in diameter, spreading branches forming a broad symmetrical head, and slender branchlets light green and villose when they first appear with long matted pale hairs, dull red-brown and pubescent in their first season, becoming glabrous the following year, and armed with occasional thin nearly straight bright chestnut-brown shining spines usually about 1½′ long.

Distribution. Rich bottom-lands of the streams of Shannon County, southern Missouri.

85. Cratægus lanuginosa Sarg.

Leaves ovate to suborbicular, acute or rounded and short-pointed at apex, broadly cuneate or rounded at the entire base, coarsely and sharply doubly serrate above with glandular teeth, and often irregularly divided above the middle into short broad acute lateral lobes, less than half grown when the flowers open during the last week of April and then dark green and villose above and covered below with a thick coat of hoary tomentum, and at maturity subcoriaceous, dark blue-green, lustrous and scabrate on the upper surface, yellow-green and tomentose on the lower surface, 1½′—2′ long, and 1′—1½′ wide, with a thick midrib, and 3—5 pairs of stout primary veins extending obliquely to the point of the lobes; petioles stout, tomentose, ½′—¾′ in length; leaves at the end of vigorous shoots often broad-ovate, very coarsely glandular-serrate, rounded or truncate at base, and frequently 3′ long and wide. Flowers ¾′ in diameter, on short stout pedicels covered with long matted pale hairs, in compact many-flowered hoary-tomentose corymbs, with large glandular-serrate conspicuous bracts and bractlets persistent until the flowers open; calyx-tube broadly obconic, hairy, the lobes short, broad, acute, glandular with minute stipitate glands, densely villose on the outer surface and slightly villose on the inner surface; stamens 20; anthers rose color; styles 5, surrounded at base by large tufts of snow-white hairs. Fruit ripening at the end of October, on short tomentose erect pedicels, in few-fruited clusters, subglobose to short-oblong, rounded and slightly hairy at the ends, ½′ in diameter; calyx enlarged, with villose coarsely serrate usually erect spreading or incurved persistent lobes bright red on the upper side near the base; flesh thin, orange color, dry and mealy; nutlets 5, thin, rounded and very irregularly ridged on the back, about ¼′ long.

A tree, sometimes 25° high, with a stout trunk covered with pale bark, spreading and erect branches, and stout zigzag branchlets light green and villose early in the season, dull red-brown and sparingly villose or pubescent at the end of their first year, becoming dark or light gray-brown, and armed with many long straight purple shining ultimately ashy gray spines 1¼′—3½′ in length.

Distribution. Southwestern Missouri; common near Webb City, Jasper County; well distinguished by the distinctly blue color of the small leaves, the dark crimson hard fruits and by the remarkable development of the spines unusual in the species of this group.

86. Cratægus arnoldiana Sarg.

Leaves broad-ovate or rarely oval, acute, regularly divided above the middle into numerous short acute lobes, and coarsely doubly serrate with straight glandular teeth except at the rounded truncate or occasionally cuneate base, coated with dense matted pale hairs when they unfold, about half grown when the flowers open at the end of May or early in June and then roughened above by stout stiff hairs and soft-pubescent below, and at maturity thin, smooth, very dark green and lustrous above, paler below, and slightly villose on the under side of the slender midrib, and of the thin prominent primary veins extending to the point of the lobes, 2′—3′ long and wide; petioles slender, densely villose early in the season, becoming puberulous, ¾′—1½′ in length; leaves at the end of vigorous shoots acute or acuminate, round or obtusely cuneate at base, more deeply lobed, often 3′—4′ long and 3′ wide. Flowers about ¾′ in diameter, on slender pedicels, in broad many-flowered tomentose corymbs; calyx-tube broadly obconic, densely tomentose, the lobes narrow, elongated, acuminate, glandular-serrate, villose on both surfaces; stamens 10; anthers, large, pale yellow; styles 3—5, usually 3 or 4, surrounded at base by a broad ring of thick hoary tomentum. Fruit ripening about the middle of August and mostly falling before the first of September, on stout pedicels, in erect spreading or rarely drooping few-fruited villose clusters, subglobose but rather longer than broad, bright crimson marked by many large pale dots, villose, particularly toward the ends, with long scattered white hairs, ¾′ long; calyx little enlarged, with elongated coarsely glandular-serrate spreading lobes often deciduous before the fruit ripens; flesh thick, bright yellow, subacid; nutlets 3 or 4, light-colored, prominently ridged on the back with a high rounded ridge, about ¼′ long.

A tree, 15°—20° high, with a short trunk 10′—12′ in diameter, stout ascending branches forming a broad open irregular head, and slender conspicuously zigzag branchlets clothed early in the season with long matted pale hairs, becoming dark orange-brown and very lustrous before midsummer, glabrous or puberulous during their first winter, bright orange-brown or gray-brown during their second year, and armed with many stout straight or slightly curved bright chestnut-brown shining spines 2½′—3′ long.

Distribution. Thickets on a dry bank in the Arnold Arboretum, valley of the Mystic River at West Medford, Middlesex County, Massachusetts, and near Lyme, New London County, Connecticut.

Often cultivated in the parks and gardens in the neighborhood of Boston; very conspicuous and easily recognized in winter by its ascending remarkably zigzag branchlets.

87. Cratægus champlainensis Sarg.

Leaves ovate, acute, rounded, truncate, slightly cordate or broad-cuneate at base, usually divided into 2 or 3 pairs of short narrow acute lobes, and coarsely often doubly serrate with glandular teeth, roughened above by short pale hairs and villose below when they unfold, nearly fully grown when the flowers open early in June, and at maturity thick and firm in texture, conspicuously blue-green and glabrous above, light yellow-green and somewhat pubescent below on the slender midrib and remote primary veins, 2′—2½′ long, and 1′—1½′ wide; petioles slender, more or less tomentose early in the season, usually becoming glabrous and light red below the middle before autumn, and ¾′—1′ in length; leaves at the end of vigorous shoots broad-ovate, rounded or slightly cordate at base, more deeply lobed, and often 3′—4′ long and wide. Flowers ¾′ in diameter, on short slender densely villose pedicels, in compact few-flowered densely villose corymbs; calyx-tube narrowly obconic, coated with thick hoary tomentum, the lobes lanceolate, finely glandular-serrate, tomentose on the outer surface usually only below the middle, villose on the inner surface; stamens 10; anthers small, light yellow; styles 5, surrounded at base by tufts of pale hairs. Fruit ripening early in September and usually remaining on the branches during the remainder of the year, on short slightly pubescent pedicels, in compact erect villose clusters, obovoid to short-oblong, bright scarlet, marked by scattered pale dots, more or less villose or pubescent toward the ends; calyx prominent, persistent, with a long tube, the lobes gradually narrowed from a broad base, acuminate, finely glandular-serrate, villose, dark red on the upper side below the middle, spreading or erect; flesh thick, yellow, dry and mealy; nutlets 5, ridged on the back with a broad ridge, /16′ long.

A tree, 15°—20° high, with a tall trunk 8′—10′ in diameter, covered with deeply fissured bark separating into thin loose plate-like scales, stout wide-spreading branches forming a broad round-topped often symmetrical head, and slender somewhat zigzag branchlets coated early in the season with hoary tomentum, soon becoming glabrous and light chestnut-brown and lustrous, and armed with straight or slightly curved chestnut-brown spines 1½′—2′ long.

Distribution. Limestone ridges; valley of the St. Lawrence River near Montreal, Province of Quebec, southward through the Champlain valley to eastern New York and westward through New York, and southern Ontario to the neighborhood of Toronto.

88. Cratægus pennsylvanica Ashe.

Leaves ovate, acuminate, rounded or abruptly cuneate at base, coarsely often doubly serrate with straight glandular teeth, and slightly divided into 3 or 4 pairs of short broad acuminate lobes, slightly tinged with red when they unfold, more than half grown when the flowers open the middle of May and then thin, dark yellow-green and roughened above by short white hairs and villose on the prominent midrib and primary veins below, and at maturity thin, dark yellow-green and scabrate on the upper surface, paler, scabrate and still somewhat villose on the midrib and veins below, 2½′—3½′ long, and 2′—2¾′ wide; petioles slender, slightly wing-margined at apex, villose through the season, occasionally glandular, 1¼′—1½′ in length; leaves at the end of vigorous shoots rounded or truncate at base, coarsely serrate, more deeply lobed, and often 4′—4½′ long and broad, with a stout midrib, prominent primary veins, a conspicuously glandular petiole, and large foliaceous lunate coarsely glandular-serrate persistent stipules. Flowers ¾′—1′ in diameter, on slender densely villose pedicels in broad lax hairy mostly 8—15-flowered corymbs; calyx-tube narrowly obconic, covered with long white hairs, the lobes long, slender, acuminate, laciniately glandular-serrate, glabrous on the outer surface, villose on the inner surface; stamens 8—12; anthers faintly tinged with pink; styles 3—5. Fruit ripening and falling early in October, on short stout drooping slightly hairy pedicels, in 4—12-fruited clusters, short-obovoid, full and rounded at apex, bright orange-red marked by small pale dots, puberulous at the ends, ¾′—1′ in diameter; calyx little enlarged, with small spreading lobes dark red on the upper side, their tips often deciduous from the ripe fruit; flesh thick, orange-yellow, somewhat acidulous, edible, sometimes made into jelly; nutlets 3—5, rounded at apex, acute at base, rounded and slightly grooved or ridged on the back, about ⅓′ long.

A tree, sometimes 30° high, with a tall trunk often 18′ in diameter, covered with dark gray scaly bark, large spreading branches forming a wide symmetrical round-topped head, and stout slightly zigzag branchlets dark orange-green and more or less tinged with red when they first appear, becoming dark chestnut-brown, marked by large dark lenticels and more or less pubescent in their first season, dark red-brown the following year, and armed with stout straight or slightly curved chestnut-brown spines 1′—1½′ long.

Distribution. Meadows in low moist soil near Pittsburgh, Allegheny County, Pennsylvania.

89. Cratægus submollis Sarg.

Leaves ovate, acute, gradually narrowed and cuneate at the nearly entire base, coarsely doubly serrate above with straight glandular teeth, and divided into 3 or 4 pairs of short acute lobes, half grown at the end of May or early in June when the flowers open and then roughened above by short stiff pale hairs and soft-pubescent below, particularly on the midrib and veins, and at maturity thin, dark yellow-green and scabrate above, pale below, 3′—3½′ long, and 2′—2½′ wide, with a thick yellow midrib and remote primary veins puberulous on the lower side; petioles stout, nearly terete, more or less winged at apex, tomentose early in the season, becoming puberulous, often bright red toward the base, 1′—2′ in length; leaves at the end of vigorous shoots broad-ovate, cuneate, rounded, truncate, or occasionally slightly cordate at base, often 4′ long and 3′—3½′ wide. Flowers 1′ in diameter, on long slender villose pedicels, in broad many-flowered tomentose corymbs; calyx-tube narrowly obconic, covered with a thick coat of long matted white hairs, the lobes gradually narrowed from a broad base, acute, glandular with large red stipitate glands, glabrous or villose on the outer surface; stamens 10; anthers small, pale yellow; styles 3—5, surrounded at base by a narrow ring of long white hairs. Fruit ripening and falling during the first half of September, on elongated slender slightly villose pedicels, in broad gracefully drooping many-fruited clusters, obovoid, bright orange-red, lustrous, marked by large scattered pale dots, puberulous toward the base, about ¾′ long; calyx much enlarged, with erect coarsely glandular-serrate persistent lobes; flesh yellow, thin, subacid, dry and mealy; nutlets usually 5, rounded and slightly ridged on the back, about ⅓′ in length.

A tree, 20°—25° high, with a tall trunk occasionally a foot in diameter, ascending or spreading ashy gray branches forming a broad handsome head, and branchlets dark green and coated with hoary tomentum when they first appear, light or dark orange-brown and slightly tomentose at midsummer, becoming glabrous, lustrous, and light red-brown or dark orange-brown, and armed with numerous thin straight or somewhat curved bright chestnut-brown shining spines 2½′—3′ in length.

Distribution. Rich damp hillsides and the borders of woods and roads; valley of the St. Lawrence River from the Isle of Orleans westward; Hull County, Province of Quebec; near Ottawa, Ontario; valley of the Penobscot River and Gerrish Island, Maine to the coast of eastern Massachusetts.

90. Cratægus Ellwangeriana Sarg.

Leaves oval, acute, rounded or broad-cuneate at the entire base, irregularly divided usually only above the middle into numerous short acute lobes, and coarsely and often doubly serrate above with straight or incurved glandular teeth, about half grown when the flowers open the middle of May, and then roughened above by short pale hairs and villose below on the slender midrib and primary veins, and at maturity thin, light green and scabrate on the upper surface, pale and nearly glabrous on the lower surface, 2½′—3½′ long, and 2′—3′ wide; petioles slender, villose early in the season, finally glabrous, 1½′—2′ in length; stipules oblong-obovate, acute, villose, coarsely glandular-serrate, ½′ long, those of the upper leaves mostly persistent until after the ripening of the fruit. Flowers 1′ in diameter, on short stout hairy pedicels, in many-flowered densely villose corymbs; calyx-tube broadly obconic, villose, the lobes long, lanceolate, glandular with small pale stalked glands, villose on both surfaces; stamens 10, sometimes 8; anthers small, rose color; styles 3—5. Fruit ripening and falling at the end of September, on slender glabrous pedicels, in drooping villose many-fruited crowded clusters, short-oblong, full and rounded at the ends, bright crimson, lustrous, covered at the ends with scattered pale hairs, 1′ long, and ½′—¾′ in diameter; calyx little enlarged, the lobes elongated, glandular-serrate above the middle, villose on the inner surface, spreading, or erect and incurved; flesh thin, yellow, juicy and acid; nutlets 3—5, thick, pale brown, deeply and often doubly and irregularly grooved on the back, ¼′—⅓′ long.

A tree, sometimes 20° high, with a tall trunk often a foot in diameter, covered with pale gray scaly bark, stout ascending branches forming a broad symmetrical head, and slender zigzag branchlets dark green and clothed at first with long matted pale hairs, becoming in their first summer light chestnut-brown and slightly villose, dark chestnut-brown and very lustrous in their second year, and armed with stout straight or somewhat curved dark chestnut-brown shining spines 1½′—2′ long.

Distribution. Western New York (common) to western Pennsylvania, and through southern Ontario to southern Michigan.

91. Cratægus Robesoniana Sarg.

Cratægus spissiflora Sarg.

Leaves oblong-ovate, acute or acuminate at apex, rounded broadly cuneate or rarely cordate at the entire base, sharply doubly serrate above with slender straight gland-tipped teeth, and deeply divided into numerous broad acute or acuminate lateral lobes, villose above and densely tomentose below when they unfold, about half grown when the flowers open at the end of May and then roughened above by short rigid white hairs and pubescent below on the midrib and veins, and at maturity dark yellow-green and scabrate on the upper surface, glabrous on the lower surface, 3′—3½′ long, and 2½′—3′ wide, with a slender midrib, and 4 or 5 pairs of prominent veins extending obliquely to the point of the lobes; petioles slender, more or less wing-margined at apex, slightly grooved, sparingly glandular, villose early in the season, becoming glabrous and rose color in the autumn, 1¼′—1½′ in length; leaves at the end of vigorous shoots cordate or rarely cuneate at base, deeply lobed, often 4′ long and 3½′ wide, with a stout conspicuous glandular petiole. Flowers ⅔′ in diameter, on short slender villose pedicels, in small very compact few, usually 4—6-flowered, thin-branched villose corymbs, with oblong-obovate acuminate glandular bracts and bractlets mostly deciduous before the flowers open; calyx-tube narrowly obconic, coated with long matted white hairs, the lobes slender, acuminate, finely glandular-serrate, glabrous on the outer surface, villose on the inner surface; stamens 10; anthers dark rose color; styles 4 or 5, surrounded at base by a narrow ring of pale tomentum. Fruit ripening at the end of September or early in October, on short reddish pubescent pedicels, in compact drooping clusters, oblong-obovoid to short-oblong, scarlet, lustrous, marked by small pale dots, about ¾′ long, and ½′ in diameter; calyx little enlarged, with spreading sharply serrate lobes often deciduous from the ripe fruit; flesh thick, yellow, dry and mealy; nutlets 4 or 5, thin, acute at the ends, rounded or only slightly grooved on the back, about ⅝′ in length.

A tree, 20°—25° high, with a trunk often 1° in diameter, covered with smooth pale gray bark, and stout spreading branches forming a round-topped head, and stout slightly zigzag dark red-brown branchlets sparingly villose early in the season, soon glabrous, bright red-brown, very lustrous and marked by small pale lenticels at the end of their first season, becoming dark gray or gray-brown the following year, and armed with few stout spreading bright chestnut-brown shining ultimately gray spines 1′—1½′ long.

Distribution. Western Massachusetts through central and western New York to the neighborhood of Toronto, southern Ontario.

92. Cratægus anomala Sarg.

Leaves ovate, acute, divided above the middle into 5 or 6 pairs of short acute or acuminate lobes, and coarsely doubly serrate with spreading glandular teeth except toward the broad-cuneate or occasionally rounded base, when they unfold conspicuously plicate, covered above with short appressed pale hairs, and villose below, especially on the slender midrib, and thin remote primary veins arching to the point of the lobes, about a third grown when the flowers open at the end of May, and at maturity membranaceous, light yellow-green, smooth and glabrous above, paler and villose below, 2½′—3′ long, and 2′—3′ wide; petioles stout, glandular on the upper side with scattered dark glands, ¾′—1′ in length; leaves at the end of vigorous shoots, rounded or truncate at base, and often 4′—4½′ long and 2½′—3′ wide. Flowers saucer-shaped, ½′ in diameter when fully expanded, on elongated slender hairy pedicels, in broad loose many-flowered villose corymbs; calyx-tube narrowly obconic, coated with long matted pale hairs, the lobes long, acuminate, coarsely glandular-serrate, pubescent on the outer surface and tomentose on the inner surface; stamens usually 10, occasionally 7 or 8; anthers large, bright red; styles 4 or 5, surrounded at base by a narrow ring of pale tomentum. Fruit ripening in October, on long slender slightly pubescent pedicels, in loose many-fruited sparingly villose clusters, obovoid to oblong, gradually narrowed to the rounded base, crimson, lustrous, marked by large pale dots, slightly villose, particularly toward the full and rounded apex, ¾′—⅞′ long, ½′—⅝′ in diameter; calyx large and prominent, with elongated acuminate lobes abruptly narrowed from a broad base, dark red on the upper side, tomentose on the lower, finely glandular-serrate, spreading or closely appressed, often deciduous before the ripening of the fruit; flesh thin, light yellow, somewhat juicy; nutlets 4 or 5, thin, prominently and irregularly ridged on the back, ¼′—/16′ long.

A bushy tree, sometimes 20° high, with a short trunk 6′ in diameter, covered with pale gray-brown scaly bark, stout ascending branches, and slender somewhat zigzag branchlets at first dark green and villose with long matted white hairs, puberulous and light orange-brown during their first season, becoming glabrous and orange-brown or bright red, and armed with numerous stout straight or slightly curved bright chestnut-brown spines 1¼′—2′ long.

Distribution. Low limestone ridges near the banks of the St. Lawrence River in the Caughnawaga Indian Reservation opposite Lachine in the Province of Quebec; western Vermont (Clarendon, Rutland County); Crown Point, Essex County, and Fort Ann, Washington County, New York.

93. Cratægus noelensis Sarg.

Leaves ovate to oval, acute, acuminate or rarely rounded at apex, acutely or broadly cuneate at base, and coarsely doubly serrate with straight teeth, covered above with short white hairs and densely villose-pubescent below when they unfold, more than half grown when the flowers open at the end of April, and at maturity dark yellow-green, smooth and glabrous on the upper surface, villose-pubescent on the lower surface, 2′—3′ long, and 1¼′—2½′ wide, with a prominent midrib and thin conspicuous primary veins; petioles slender, slightly wing-margined at apex, hoary-tomentose early in the season, becoming glabrous, 1′—1¼′ in length; leaves at the end of vigorous shoots ovate, acuminate, rounded or cuneate at the broad base, more coarsely serrate, usually laterally lobed with short broad acuminate lobes, 3′—4′ long, and 2½′—3′ wide. Flowers ¾′ to nearly 1′ in diameter, on short pedicels densely covered like the narrow obconic calyx-tube and the compact 5—10-flowered corymb with long matted white hairs; calyx-lobes slender, long-acuminate, minutely glandular-serrate, slightly villose; stamens 5—10, usually 10; anthers rose color; styles 3—5, surrounded at base by a broad ring of pale tomentum. Fruit ripening in September, on slender drooping pubescent pedicels, subglobose, orange-red, ½′—⅔′ in diameter, the calyx prominent with a short tube and spreading closely appressed lobes; flesh thin, soft and yellow; nutlets 3—5, rounded at base, narrowed and rounded at apex, slightly grooved on the back, about ¼′ long.

A tree, 15°—18° high, with a trunk sometimes 1° in diameter, spreading branches forming a broad flat or round-topped head, and stout zigzag branchlets coated when they first appear with matted white hairs, reddish brown, pubescent or puberulous during their first season and gray the following year, and armed with few or many slender straight purple lustrous spines 1′—2½′ in length, sometimes persistent and compound on old trunks.

Distribution. Rich alluvial soil; in the neighborhood of Noel, McDonald County, Missouri; common.

IX. Coccineæ.

Flabellatæ Sarg.

Conspectus of the Arborescent Species.

Stamens 20; leaves yellow-green and scabrate above.

Leaves ovate; anthers deep rose-purple; fruit obovoid to short-oblong, bright red, often slightly pruinose.

94. C. neo-londinensis (A).

Leaves oblong-ovate; anthers pink; fruit obovoid, crimson, lustrous.

95. C. Hillii (A).

Stamens 10—20, usually 10; anthers pinkish purple, leaves broad-ovate, dull dark green and scabrate above; fruit short-oblong to slightly obovoid, dull red or crimson.

96. C. assurgens (A).

Stamens usually 10.

Fruit on short stout pedicels; leaves yellow-green and glabrous above.

Leaves oval, drooping, conspicuously concave; anthers purple; fruit short-oblong, dark dull red, villose at the ends.

97. C. Pringlei (A).

Leaves oval to oblong-ovate; anthers dark reddish purple; fruit short-oblong, crimson, lustrous.

98. C. lobulata (A).

Fruit on long slender pedicels; leaves broad-ovate to obovate or rhombic, dark rich green and scabrate above; anthers rose color; fruit short-oblong, bright scarlet, lustrous.

99. C. pedicellata (A).

Stamens usually 5—7, rarely 10.

Fruit obovoid to ellipsoidal; leaves oval or ovate, conspicuously yellow-green; anthers dark reddish purple; fruit crimson, lustrous.

100. C. Holmesiana (A).

Fruit short-oblong; leaves oblong-ovate, deep yellow-green, nearly smooth above; anthers pink; fruit yellowish red, glaucous.

101. C. acclivis (A).

Fruit subglobose to obovoid.

Leaves glabrous above; anthers dark rose color.

Leaves broad-ovate, thin, light yellow-green and lustrous above; fruit bright red or scarlet.

102. C. delecta (A).

Leaves oblong-ovate, subcoriaceous, dark dull green; fruit bright cherry-red, pruinose.

103. C. Eamesii (A).

Leaves scabrate above, oblong-ovate, thin, dark yellow-green; anthers pale rose color; fruit crimson.

104. C. sertata (A).

94. Cratægus neo-londinensis Sarg.

Leaves ovate, acute or acuminate, rounded, truncate or broadly concave-cuneate at the wide entire or glandular base, sharply often doubly serrate above with straight glandular teeth, and divided into numerous short narrow acuminate lateral lobes, about half grown when the flowers open the middle of May and then very thin, light yellow-green and roughened above by short white rigid hairs and paler and sparingly hairy below, and at maturity membranaceous, dull yellow-green and scabrate on the upper surface, pale green and glabrous below, or occasionally slightly hairy on the under side of the stout yellow midrib, and of the thin remote primary veins arching to the point of the lobes, 3′—4′ long, and 2½′—3½′ wide; petioles slender, nearly terete, glandular, at first slightly hairy, becoming glabrous and purplish toward the base, 1′—2′ in length; leaves at the end of vigorous shoots only slightly larger. Flowers 1′—1⅛′ in diameter, on slender sparingly villose pedicels, in lax slightly drooping usually 5—12-flowered villose or nearly glabrous corymbs, with linear often slightly falcate glandular bracts and bractlets, persistent until after the flowers open; calyx-tube narrowly obconic, covered with short matted pale hairs, the lobes gradually narrowed from a broad base, acuminate, coarsely glandular-serrate below the middle, glabrous on the outer, villose on the inner surface; stamens 17—21, usually 20; anthers deep rose-purple; styles 4 or 5, usually 5, surrounded at base by a narrow ring of pale tomentum. Fruit ripening and beginning to fall early in September, on stout villose or glabrous pedicels, in large drooping few-fruited clusters, obovoid or short-oblong, bright red, often slightly pruinose, marked by numerous minute pale dots, ⅝′—¾′ long, ½′—⅝′ in diameter; calyx enlarged, prominent, with spreading or erect and incurved coarsely serrate persistent lobes, their upper surface bright red below the middle and covered above with soft white hairs; flesh thick, orange-yellow, soft, juicy and acidulous; nutlets 4 or 5, thin, narrowed at the ends, acute at base, rounded at apex, rounded and sometimes broadly grooved on the back, about /16′ long.

A tree, often 20° high, with a tall trunk 8′—10′ in diameter, covered with light grayish brown slightly fissured bark, large spreading and drooping branches forming an open head often 20° across, and slender branchlets olive-green and slightly hairy when they first appear, dull red-brown and marked by many large pale lenticels during their first season, becoming light gray and rather lustrous, and armed with stout straight dark purple shining ultimately gray spines often 2′ long.

Distribution. Borders of woods near the shores of Fisher’s Island Sound, Mumford’s Point, Groton, and near Lyme, New London County, Connecticut.

95. Cratægus Hillii Sarg.

Leaves oblong-ovate, acuminate, rounded or rarely cuneate at the broad entire base, coarsely doubly serrate above with straight glandular teeth, and divided into numerous short acuminate lateral lobes, when they unfold coated above with short lustrous white hairs and densely tomentose below, particularly on the midrib and veins, about one fourth grown when the flowers open the middle of May and then roughened above by short hairs and villose below, and at maturity thin, light yellow-green and scabrate on the upper surface, pale yellow-green on the lower surface, 2½′—3′ long, and 2′—2½′ wide, with a slender midrib often slightly hairy near the base, and 4 or 5 pairs of thin primary veins extending obliquely to the point of the lobes; petioles slender, densely villose early in the season, slightly hairy in the autumn, and ⅝′—1½′ in length; leaves at the end of vigorous shoots often truncate or slightly cordate at base, deeply lobed with broad triangular lobes, and 3½′—4′ long and wide, with a stout rose-colored glandular petiole, and hairy lunate glandular-serrate stipules. Flowers about ¾′ in diameter, on slender densely villose pedicels, in broad many-flowered hairy compound corymbs, their large linear to oblong bracts and bractlets occasionally persistent until midsummer; calyx-tube narrowly obconic, thickly covered with long spreading white hairs, the lobes abruptly narrowed at base, broad, acuminate, coarsely glandular-serrate, glabrous on the outer surface, villose on the inner surface; stamens 20; anthers pink; styles 4 or 5, surrounded at base by a narrow ring of pale tomentum. Fruit ripening from the middle to the end of September, on slender puberulous pedicels, in drooping few-fruited clusters, obovoid, broad and rounded at apex, gradually narrowed to the rounded base, crimson, lustrous, marked by small pale dots, ½′—⅝′ long, ⅜′—½′ in diameter; calyx only slightly enlarged, with closely appressed coarsely serrate lobes often deciduous from the ripe fruit; flesh yellow, thin, acidulous, juicy; nutlets 4 or 5, thin, gradually narrowed and acute at the ends, irregularly ridged and sometimes grooved on the back, about ⅜′ long.

A tree, 25°—30° high, with a trunk sometimes a foot in diameter and 6° or 7° long, covered with close light gray bark tinged with red and divided by shallow fissures into small plates, stout ascending branches forming an open irregular often round-topped head, and slender nearly straight branchlets densely villose when they first appear, dark orange color tinged with red and sparingly villose when the flowers open, becoming bright red-brown and lustrous at the end of their first season and dark dull reddish brown the following year, and sparingly armed with slender nearly straight red-brown shining spines 1½′—2′ long.

Distribution. Open woods near the borders of streams in moist rich soil; northeastern Illinois (Thatcher’s Park, Glendon Park, and River Forest, Cook County); not common.

96. Cratægus assurgens Sarg.

Leaves broad-ovate, acuminate, rounded or rarely cuneate at the wide entire base, sharply doubly serrate above with straight gland-tipped teeth, and slightly divided, into 3 or 4 pairs of small acuminate lobes, about one third grown when the flowers open the middle of May and then roughened above by short white hairs and glabrous or sparingly villose below, with persistent hairs on the slender yellow midrib, and on the veins arching obliquely to the point of the lobes, and at maturity membranaceous, dull dark green and scabrate on the upper surface, light yellow-green on the lower surface, 2¾′—3½′ long, and 2¼′—2¾′ wide; petioles slender, villose early in the season, becoming pubescent, 1′—1½′ in length; leaves at the end of vigorous shoots often deeply lobed, coarsely serrate, sometimes 4′ long and wide, with long stout glandular petioles and foliaceous lunate acuminate coarsely glandular-serrate persistent stipules. Flowers ¾′—⅝′ in diameter, on short villose pedicels, in compact 8—15-flowered hairy corymbs, with oblong, acuminate, glandular bracts and bractlets, deciduous with the opening of the flowers; calyx-tube narrowly obconic, sparingly villose, the lobes long, narrow, acuminate, tipped with minute red glands, finely glandular-serrate, glabrous on the outer, pubescent on the inner surface; stamens 10—20, usually 10; anthers pinkish purple; styles 4 or 5, surrounded at base by tufts of pale hairs. Fruit ripening from the 15th to the 20th of September, and usually falling about the 1st of October, on short glabrous pedicels, in drooping few-fruited clusters, short-oblong to slightly obovoid, dull red to crimson, ½′—⅝′ long, about ½′ wide; calyx sessile, with spreading closely appressed serrate usually persistent lobes; flesh thin, pale yellow or nearly white, acidulous; nutlets 4 or 5, broad, narrow and acute at the ends, prominently ridged on the back with a high narrow ridge, or often grooved, about ¼′ long.

A tree, sometimes 25° high, with a trunk 2′—6′ in diameter and often 6°—9° long, covered with close dark gray bark, ascending branches forming an oblong, open head, and slender branchlets light orange-yellow and covered when they first appear with long scattered caducous white hairs, becoming bright red-brown and lustrous, and dark gray-brown the following year, and armed with many stout usually slightly curved bright red-brown shining spines, 1′—1½′ long.

Distribution. River banks and low woods in rich soil; northeastern Illinois (Leyden township, La Grange, Thatcher’s Park, Cook County, Highland Park, Deerfield, Wauconda, Lake County); Fox Point, Milwaukee County, Wisconsin.

97. Cratægus Pringlei Sarg.

Leaves oval, acute, rounded or often abruptly narrowed and cuneate at base, occasionally irregularly lobed above the middle with short broad acute lobes, and coarsely and often doubly serrate with glandular teeth, as they unfold villose on both surfaces, and often more or less tinged with red, when the flowers open, usually in the last week of May, roughened above by short closely appressed pale hairs and glabrous below with the exception of a few hairs on the slender midrib and remote primary veins, and at maturity thin, glabrous, and bright yellow-green on the upper surface, pale below, 2′—2½′ long, and 1¾′—2¼′ wide, usually conspicuously concave by the gradual turning down of the blades from the midrib to the margins, drooping on long thin slender glandular petioles at first villose, ultimately glabrous, 1′—1¾′ in length; leaves at the end of vigorous shoots sometimes truncate or slightly cordate at the base, and frequently 3′ long and wide. Flowers about ¾′ in diameter, on stout hairy pedicels, in many-flowered compound villose corymbs; calyx-tube narrowly obconic, villose, particularly toward the base, the lobes narrow, acuminate, coarsely glandular-serrate, villose on both surfaces or only on the inner surface; stamens 10, occasionally 5—10; anthers small, purple; styles 3—5, surrounded at the base by conspicuous tufts of pale tomentum. Fruit ripening and falling late in September or early in October, on stout pedicels, in erect villose mostly few-fruited clusters, short-oblong, dark dull red, marked by few dark dots, villose at the ends with long scattered pale hairs, ¾′ long and ⅝′ in diameter; calyx little enlarged, the lobes gradually narrowed from a broad base, acuminate, glandular-serrate, often erect; flesh thick, yellow, dry and acid, with a disagreeable flavor; nutlets 3—5, rounded and slightly ridged on the back, ⅓′ long.

A tree, occasionally 25° high, with a tall trunk 10′—12′ in diameter, covered with thin bark separating into large flakes broken into small loose dark red-brown scales, stout branches forming a wide symmetrical head, and slightly zigzag branchlets at first dark green and villose, soon becoming glabrous, chestnut-brown and lustrous, bright orange-brown during their second year, and armed with thick straight or somewhat curved chestnut-brown spines often 1½′ long.

Distribution. Southern New Hampshire, through southern Vermont to western Massachusetts and eastern New York; through central and western New York and southern Ontario to northeastern Ohio (Plymouth, Ashtabula County), the southern peninsula of Michigan and northeastern Illinois.

98. Cratægus lobulata Sarg. Red Haw.

Leaves oval to oblong-ovate, acute at apex, broad-cuneate or rounded at the entire base, sharply and often doubly serrate above with straight glandular teeth, and deeply divided into numerous narrow acute or acuminate lobes spreading or pointing to the apex or to the base of the leaf, when they first appear and until after the opening of the flowers during the last week in May covered above with short soft pale hairs and slightly pubescent below on the slender midrib, and thin primary veins arching to the point of the lobes, and at maturity thin, dark yellow-green and glabrous on the upper surface, paler on the lower surface, with occasional short white hairs toward the base of the midrib, 2½′—3½′ long and 2′—2½′ wide; petioles slender, nearly terete, at first tomentose, particularly at the base, becoming pubescent or nearly glabrous and bright red, 1′—1½′ in length; leaves at the end of vigorous shoots broad-ovate, rounded or truncate at the broad base, divided into numerous acuminate lateral lobes, often 3½′—4′ long and 3′—3½′ wide. Flowers ¾′ in diameter, on elongated slender pedicels, in rather compact many-flowered tomentose corymbs, with linear-lanceolate glandular-serrate bright red bracts and bractlets; calyx-tube broadly obconic, glabrous, or villose toward the base, dark red, the lobes gradually narrowed from a broad base, glabrous, coarsely glandular-serrate with large dark red stipitate glands; stamens usually 10, occasionally 5—10; anthers small, dark reddish purple; styles 3—5, sometimes surrounded at the base by a ring of pale tomentum. Fruit ripening and falling early in October, on short stout pedicels, in erect compact tomentulose clusters, short-oblong, somewhat flattened at the rounded ends, bright crimson, very lustrous, marked by occasional small white dots, about ¾′ long and ⅝′ in diameter; calyx little enlarged, the lobes small, lanceolate, coarsely glandular-serrate, tomentose on the upper surface, erect and incurved, persistent; flesh thick, yellow, sweet and juicy; nutlets 3—5, thin, dark colored, ridged and often grooved on the back, ¼′ long.

A tree, occasionally 35° high, with a straight trunk often a foot in diameter, covered with dark red-brown fissured bark broken into small thick plate-like scales, stout generally ascending branches forming an open usually narrow irregular head, and slender branchlets, dark green and covered with matted pale hairs when they first appear, becoming bright chestnut-brown and very lustrous during their first season, and light orange-brown the following year, and armed with many stout nearly straight chestnut-brown spines rarely more than 1′ in length.

Distribution. Burlington, Chittenden County, Vermont, and southward through the Champlain valley to Crown Point, Essex County and to the neighborhood of Albany, New York; western Massachusetts to southern Connecticut (Stratford, Fairfield County); common.

99. Cratægus pedicellata Sarg.

Leaves broad-ovate or occasionally obovate or rhombic, acute or acuminate, broad-cuneate or rounded at the entire base, coarsely often doubly serrate above with spreading glandular teeth, and divided above the middle into 4 or 5 pairs of short acute or acuminate lobes, nearly two thirds grown when the flowers open during the last week in May, and then roughened above by short rigid pale hairs and glabrous below, and at maturity membranaceous, dark rich green and scabrate on the upper surface, pale on the lower surface, 3′—4′ long, and 2′—3′ wide, with a slender midrib, and thin remote primary veins arching to the point of the lobes; petioles slender, nearly terete, glandular with minute scattered dark glands, at first villose, becoming glabrous, 1½′—2½′ in length; leaves at the end of vigorous shoots sometimes truncate or slightly cordate at base, more deeply lobed, often 3′—4′ long and 3′ wide. Flowers ½′ in diameter, on long thin pedicels, in loose lax many-flowered slightly villose corymbs; calyx-tube narrowly obconic, glabrous, the lobes broad, acute, very coarsely glandular-serrate; stamens usually 10; anthers rose color; styles 5, surrounded at base by a conspicuous ring of pale tomentum. Fruit ripening and falling during September, on long slender pedicels, in few-fruited drooping glabrous clusters, obovoid until nearly fully grown, becoming short-oblong when fully ripe, rounded at the ends, bright scarlet, lustrous, marked by numerous small dark dots, ¾′ long, and ½′—⅝′ in diameter; calyx large and conspicuous, the lobes much enlarged, coarsely serrate, and usually erect and incurved; flesh pale, thin, dry and mealy; nutlets 5, narrowed and acute at the ends, rounded and deeply grooved on the back, about ⅓′ long.

A tree, 18°—20° high, with a tall trunk sometimes a foot in diameter, covered with close red-brown scaly bark, long comparatively slender spreading or ascending branches forming a handsome symmetrical head, and thin branchlets dark chestnut-brown and slightly villose at first, becoming very lustrous and ashy gray in their second year, and armed with straight or slightly curved shining chestnut-brown spines 1½′—2′ long.

Distribution. Central and western New York to western Pennsylvania (Allegheny and Crawford counties), and to southern Ontario to the neighborhood of Toronto and London; common; passing into var. gloriosa Sarg. differing in its rather larger flowers with pink anthers, larger and more lustrous fruit often mammillate at base and ripening a few days earlier and in its convex leaves. A tree, 20°—25° high, with a trunk often 1° in diameter, and a symmetrical round-topped head; Rochester, Munroe County, New York; not common.

100. Cratægus Holmesiana Ashe.

Leaves oval or ovate, acute or acuminate at apex, rounded or broad-cuneate at base, coarsely doubly serrate above the middle with straight teeth tipped at first with prominent dark red caducous glands, and usually divided into 3 or 4 pairs of short acute or acuminate lateral lobes, when they unfold dark red, roughened by rigid pale hairs on the upper surface, and glabrous or sometimes villose on the lower surface, scabrate above, pale yellow-green and nearly half grown when the flowers open early in May, and at maturity thick and firm, almost smooth, conspicuously yellow-green, usually about 2′ long and 1¾′ wide, with a prominent midrib often bright red on the lower side toward the base, and 4—6 pairs of slender primary veins arching to the point of the lobes; petioles slender, nearly terete, glandular, glabrous or sometimes puberulous while young, 1′—1½′ in length; leaves at the end of vigorous shoots often broad-ovate to oval, rounded, truncate or slightly cordate at base, more coarsely serrate and more deeply lobed, and frequently 4′ long and 3′ wide. Flowers cup-shaped, ½′—¾′ in diameter, on long slender glabrous pedicels, in loose glabrous or rarely puberulous many-flowered corymbs, with oblanceolate or linear acute glandular caducous bracts and bractlets; calyx-tube narrowly obconic, glabrous, more or less deeply tinged with red, the lobes long, acuminate, glandular-serrate, or often nearly entire; glabrous on the outer surface, villose-pubescent on the inner surface; stamens usually 5, sometimes 6—8; anthers large, dark reddish purple; styles usually 3, surrounded at base by a narrow ring of pale tomentum. Fruit ripening and falling early in September, on long slender pedicels, in many-fruited drooping clusters, obovoid to ellipsoidal, crimson, lustrous, marked by occasional small dark dots, about ½′ long, and ⅓′ in diameter; calyx enlarged, conspicuous, with erect and incurved glandular-serrate lobes, bright red toward the base on the upper side; flesh thin, yellow, dry and mealy, with a disagreeable flavor; nutlets usually 3, light chestnut-brown, prominently grooved and ridged on the back with a broad rounded ridge, about ¼′ long.

A tree, often 30° high, with a tall straight trunk 10′—15′ in diameter, covered with pale gray-brown or nearly white scaly bark, stout ascending branches forming an open irregular rather compact head, and stout glabrous branchlets dark green more or less tinged with red when they first appear, becoming bright chestnut-brown or orange-brown and lustrous, and ultimately ashy gray, and armed with occasional thick mostly straight bright chestnut-brown shining spines 1½′—2′ long.

Distribution. Rich moist hillsides and the borders of streams and swamps, neighborhood of Montreal and southern Ontario to the coast of southern Maine, central and western Massachusetts, Rhode Island, western New York, and eastern Pennsylvania; most abundant and of its largest size on the hills of Worcester County, Massachusetts. In Sellersville, Bucks County, Pennsylvania, in a form of this species (var. villipes Ashe) the young branchlets, petioles, and corymbs are often puberulous and the under surface of the leaves more or less hairy, especially on the midrib and veins. Passing into var. tardipes Sarg. differing from the type in its darker green leaves somewhat rougher on the upper surface, flowers often ⅘′ in diameter on villose pedicels, and in the shorter slightly hairy pedicels of the fruit ripening early in October.

A tree, in size, habit and bark similar to the species; southern Ontario (neighborhood of Toronto, common, near London, bank of the St. Claire River below Sarnia and Walpole Island, Lamberton County); Province of Quebec (Montreal, Caughnawaga, Isle Perrot, St. Ann’s and Hemmingford); central and western New York.

101. Cratægus acclivis Sarg.

Leaves oblong-ovate, acuminate, broad-cuneate or rounded at the entire base, coarsely doubly serrate above with straight gland-tipped teeth, and deeply divided into numerous wide-spreading acuminate lateral lobes, when they unfold tinged with red, densely villose on the upper surface, pubescent on the midrib and veins below, about half grown when the flowers open during the last week of May and then light yellow-green, slightly roughened above by short white hairs and pubescent on the midrib and veins below, and at maturity membranaceous, dark yellow-green and nearly smooth above, pale yellow-green and glabrous below, 2½′—3′ long, and 2′—2½′ wide, with a stout yellow midrib, and 5 or 6 pairs of primary veins extending obliquely to the point of the lobes; petioles slender, slightly wing-margined at apex, glandular with numerous small dark glands, densely villose early in the season, becoming puberulous or glabrous in the autumn, 1½′—2′ in length; leaves at the end of vigorous shoots broad-ovate, acuminate, cordate at the wide base, deeply divided into wide acute lateral lobes, and often 4′—5′ long and wide, with foliaceous, lunate, coarsely glandular-serrate stipules, 1½′ wide, and persistent throughout the season. Flowers ¾′ in diameter, on slender densely villose pedicels, in broad lax many-flowered long-branched hairy corymbs, their bracts lanceolate, glandular, large and conspicuous, persistent until after the flowers open; calyx-tube narrowly obconic, covered with a thick coat of long matted hairs, the lobes long slender, acuminate, serrate with occasional large gland-tipped teeth, glabrous on the outer surface, slightly villose on the inner surface; stamens usually 5; anthers pink; styles mostly 5. Fruit ripening the middle of September and soon falling, on long slender slightly hairy pedicels, in many-fruited drooping clusters, short-oblong, broad and rounded at the ends, yellowish red, glaucous, marked by occasional pale dots, about ¾′ long and ⅝′ wide; calyx sessile, with usually erect enlarged coarsely serrate lobes villose on the upper side and often deciduous from the ripe fruit; flesh thick, yellow, rather juicy; nutlets usually 5, narrow and acute at the ends, ridged with a high broad ridge, or rounded and slightly grooved on the back, about ⅝′ long.

A tree, 25°—30° high, with a short trunk occasionally 4′—5′ in diameter, covered with smooth light gray bark, numerous erect branches forming an oblong open very irregular head, and stout slightly zigzag branchlets coated when they first appear with long matted pale hairs, light red-brown and lustrous, marked by small pale lenticels, and pubescent at the end of their first season, becoming dull red or orange-brown the following year, and armed with stout straight or curved bright red-brown shining spines 1¼′—2′ long.

Distribution. New York: near Albany, Albany County, steep banks of the gorge of the Genesee River, Rochester, Munroe County, banks of the Niagara River, Niagara Falls, Niagara County, and near Buffalo, Erie County; common.

102. Cratægus delecta Sarg.

Leaves broad-ovate, acute or acuminate at apex, rounded or broad-cuneate at the entire base, sharply often doubly serrate above with straight glandular teeth, and divided usually only above the middle into numerous short acuminate lateral lobes, when they unfold tinged with red and covered with glistening white hairs more abundant below than above, nearly half grown when the flowers open during the first half of May and then roughened on the upper surface by short white hairs and glabrous or sparingly villose on the midrib and veins below with scattered hairs sometimes persistent through the season, and at maturity membranaceous, light yellow-green, lustrous and glabrous above, paler below, 1½′—2′ long and wide, with a stout yellow midrib, and 6 or 7 pairs of slender primary veins arching obliquely to the point of the lobes; turning purplish in the autumn before falling; petioles slender, covered early in the season with matted pale hairs, becoming glabrous, slightly glandular, often tinged with red below the middle, ¾′—1′ in length; leaves at the end of vigorous shoots sometimes long-pointed at apex and slightly cordate at base, more deeply lobed and more coarsely serrate, and often 3′—4′ long and wide. Flowers ¾′—1′ in diameter, on long slender slightly hairy pedicels, in broad villose 10—15-flowered sparingly villose corymbs, with glandular caducous bracts and bractlets; calyx-tube broadly obconic, villose or nearly glabrous, the lobes acuminate, coarsely glandular-serrate, glabrous on the outer surface, villose on the inner surface; stamens 5—10, usually 5; anthers dark rose color; styles 3—5, usually 5. Fruit ripening from the first to the middle of September and soon falling, on stout glabrous pedicels, in drooping few-fruited clusters, subglobose to slightly obovoid, bright red or scarlet, becoming purple when fully ripe, ½′—¾′ long, and ⅔′—¾′ in diameter; calyx prominent, with erect and incurved coarsely serrate lobes; flesh thick, yellow, juicy, mildly acid and edible; nutlets 3—5, usually 5, narrowed and acute at the ends, rounded and very irregularly ridged on the back, ¼′—/16′ long.

A tree, sometimes 30° high, with a trunk rarely 1° in diameter and 6°—9° long, covered with light gray slightly fissured smooth bark, spreading or ascending branches forming an oblong open head, and slender branchlets at first slightly villose, becoming glabrous, dull red, and ultimately gray or olive-gray, and armed with stout nearly straight spines much thickened below the middle, dark chestnut-brown and lustrous, becoming dull brown or gray, and usually 1′—2′ long.

Distribution. Pastures, open woods or their borders; northeastern Illinois (Lockport, Will County, Wauconda, Fort Sheridan, Deerfield, Lake Forest, Highland Park, Lake County).

103. Cratægus Eamesii Sarg.

Leaves oblong-ovate, acute or acuminate, concave-cuneate or rounded at the entire or glandular base, sharply often doubly serrate above with straight glandular teeth, and divided into numerous short acute lateral lobes, about half grown when the flowers open the middle of May, and then membranaceous, light yellow-green and roughened above by short rigid white hairs and pale and glabrous below with the exception of a few hairs on the midrib, and slender primary veins arching to the point of the lobes, and at maturity subcoriaceous, glabrous, dark rather dull green and smooth above, pale yellow-green below, 3′—3½′ long, and 2′—2½′ wide; petioles slender, wing-margined above, villose at first, becoming glabrous, 1′—1½′ in length; leaves at the end of vigorous shoots usually rounded or truncate at the broad base, more deeply lobed, often 3½′—4′ long and 3½′ wide. Flowers about ¾′ in diameter, on slender slightly hairy pedicels, in crowded compact 5—25, usually 15—18-flowered sparingly villose corymbs, with linear-obovate coarsely glandular reddish bracts and bractlets, mostly deciduous before the flowers open; calyx narrowly obconic, glabrous, the lobes long, slender, glandular with large bright red stipitate glands, glabrous on the outer, slightly villose on the inner surface; stamens 5—10, usually 5—8; anthers deep rose-purple; styles 4 or 5, surrounded at base by a narrow ring of pale pubescence. Fruit ripening early in September and soon falling, on stout glabrous pedicels, in large many-fruited drooping clusters, short-oblong to slightly ovoid, rounded at the ends, bright cherry-red, lustrous, pruinose, marked by few large dark dots, ⅝′—¾′ long, and about ½′ in diameter; calyx only slightly enlarged, the lobes erect and incurved, coarsely serrate, dark red on the upper side below the middle, their tips deciduous from the ripe fruit; flesh thick, pale yellow, juicy; nutlets 4 or 5, narrow at the ends, irregularly ridged often with a high broad ridge, and sometimes grooved on the back, about ¼′ long.

A tree, occasionally 20° high, with a trunk a foot in diameter, ascending branches forming a narrow open head, and stout glabrous branchlets bright reddish brown and rather lustrous during their first season, becoming light gray slightly tinged with red in their second year, and armed with stout straight or slightly curved spines 1′—1½′ long; or occasionally shrubby, with a short trunk divided near the ground into several spreading stems.

Distribution. Rich moist ground, Stratford, Fairfield County (E. H. Eames), and Ansonia, New Haven County, Connecticut (E. B. Harger).

104. Cratægus sertata Sarg.

Leaves oblong-ovate, acuminate, rounded, truncate, subcordate or rarely cuneate at the broad base, finely and often doubly serrate with straight gland-tipped teeth, and deeply divided into 5 or 6 pairs of wide acuminate lobes, when they unfold coated above with short pale hairs and villose below on the midrib and veins, about half grown and villose when the flowers open during the first half of May, and at maturity membranaceous, dark yellow-green and scabrate on the upper surface, pale yellow-green and glabrous on the lower surface, 2½′—3′ long, and 1½′—2′ wide, with a thin yellow midrib, and slender primary veins arching obliquely to the point of the lobes; petioles slender, slightly grooved, villose early in the season, ultimately glabrous, sparingly glandular, 1½′—3′ in length; leaves at the end of vigorous shoots broad-ovate, rounded or slightly cordate at base, often 3′ long and 2½′ wide. Flowers ¾′—1′ in diameter, on slender pedicels, in broad 10—15-flowered densely villose corymbs, with linear to linear-obovate glandular large and conspicuous caducous bracts and bractlets; calyx-tube broadly obconic, glabrous above, villose below, the lobes abruptly narrowed from the base, broad, acuminate, tipped with small red glands, coarsely glandular-serrate, glabrate on the outer surface, pubescent on the inner surface; stamens 5—10, usually 5; anthers pale rose color; styles 3—5, surrounded at base by tufts of pale hairs. Fruit ripening about the middle of September and soon falling, on slender villose or pubescent pedicels, in drooping many-fruited clusters, subglobose to slightly obovoid, rounded at the ends, bright red and lustrous, becoming darker or crimson when fully ripe, marked by occasional large pale dots, about ½′ long and wide; calyx prominent, with enlarged mostly erect incurved serrate lobes; flesh thin, yellow, aromatic, pleasantly acid; nutlets 3—5, usually 4, thin, narrow and acute at the ends, slightly ridged on the back with a wide or narrow ridge, ⅜′ long.

A tree, 10°—20° high, with a trunk 6′—8′ in diameter and often 4°—5° long, covered with close dark gray bark separating into long narrow thin plate-like scales, stout spreading branches forming a handsome open head, and slender nearly straight branchlets thickly coated when they first appear with matted pale hairs, light brown and lustrous at the end of their first season, and dark gray-brown the following year, and unarmed or armed with stout nearly straight or curved spines 1′—2½′ long.

Distribution. Open woods and pastures in rich moist soil; northeastern Illinois (Mokena, Will County, Glenellyn, Dupage County, Barrington, Glendon Park, Cook County, Highland Park, Lake Zurich, Lake County); Fox Point, Milwaukee County, Wisconsin.

X. Dilatatæ.

Conspectus of the Arborescent Species.

Flowers in broad 6—12-flowered corymbs.

Leaves broad-ovate; fruit bright scarlet.

105. C. dilatata (A).

Leaves nearly orbicular to oval; fruit dull red blotched with green, or orange-red.

106. C. suborbiculata (A).

Leaves ovate to slightly obovate; fruit crimson, pruinose.

107. C. hudsonica (A).

Flowers in very compact 5—7-flowered corymbs; leaves broad-ovate; fruit usually broader than high, much flattened at the ends, dark crimson, very lustrous.

108. C. coccinioides (A).

105. Cratægus dilatata Sarg.

Leaves broad-ovate, acute, truncate, cordate or slightly rounded at the broad base, coarsely and generally doubly and irregularly serrate above with straight teeth tipped with large dark glands, unequally lobed usually with 2 or 3 pairs of acute or acuminate lateral lobes, about one third grown when the flowers open at the end of May, and then light yellow-green, conspicuously plicate, roughened on the upper surface with short stiff white hairs and glabrous on the lower surface, and at maturity smooth and glabrous, dark green above, pale below, 2′—2½′ long, and almost as wide as long, with a slender midrib and 4 or 5 pairs of thin primary veins; petioles slender, somewhat glandular, at first villose, soon glabrous, often dark red toward the base after midsummer, 1′—2′ in length; leaves at the end of vigorous shoots often 4′—5′ long, and frequently rather broader than long. Flowers 1′—1⅛′ in diameter, on slender elongated hairy pedicels, in broad, loose, usually 8—12-flowered slightly villose corymbs, with lanceolate bracts and bractlets glandular like the inner bud-scales with dark red glands; calyx-tube broadly obconic, covered toward the base with matted pale hairs, nearly glabrous above, the lobes broad, acuminate, coarsely glandular with large scattered red glands, glabrous on the outer surface and generally slightly villose on the inner surface; stamens 20; anthers large, rose color; styles usually 5, surrounded at base by small tufts of white hairs. Fruit ripening and falling early in September, on slender pedicels, in many-fruited drooping clusters, subglobose, bright scarlet, marked by numerous small dark dots, about ¾′ in diameter; the calyx much enlarged, with spreading coarsely serrate lobes bright red on the upper side toward the base; flesh thin, sweet and yellow; nutlets 5, thin, rounded and prominently ridged on the back, about ¼′ long.

A tree, occasionally 20° high, with a tall straight trunk, covered with light gray-brown scaly bark, branches spreading into a wide round-topped symmetrical head, and short glabrous slightly zigzag branchlets armed with few stout straight light brown shining spines 1′—2′ long.

Distribution. Eastern Massachusetts, coast of Rhode Island, western Vermont, in the neighborhood of Albany, New York, and near Montreal, Province of Quebec.

106. Cratægus suborbiculata Sarg.

Leaves nearly orbicular to oval or rarely to oblong, short-pointed at apex, broad and rounded or broad-cuneate at the entire base, sharply doubly serrate above with slender straight or incurved glandular teeth, and often divided above the middle into 3 or 4 pairs of short acute lobes, when they unfold pale yellow-green and somewhat villose on the upper surface toward the base and below in the axils of the principal veins, about a third grown when the flowers open during the first week of June, and at maturity thin and firm in texture, dull dark green above, paler below, usually about 1½′ long and broad, with a slender midrib and 4 or 5 pairs of thin primary veins; petioles slender, slightly glandular, more or less winged above, ⅝′—1′ in length; leaves at the end of vigorous shoots nearly orbicular to oval, more coarsely serrate and more deeply lobed, and frequently 3′ long and wide, their petioles often broadly winged and conspicuously glandular. Flowers ¾′ in diameter, on short stout pedicels, in compact 6—12-flowered glabrous corymbs; calyx broadly obconic, the lobes gradually narrowed from a broad base, long, acuminate, entire or occasionally obscurely denticulate; stamens 20; anthers small, rose color; styles 5, surrounded at base by a broad ring of hoary tomentum. Fruit falling in October without becoming mellow, on short rigid pedicels, in few-fruited erect clusters, subglobose, often rather longer than broad, about ⅝′ in diameter, dull red more or less blotched with green, or often wholly green on one face, or scarlet in one form; calyx enlarged, prominent, with a broad deep cavity and nearly entire wide-spreading lobes; flesh yellow, thin, dry and hard; nutlets 5, broad and thick, narrow and rounded at the ends, obscurely and unequally grooved on the back, about ¼′ long.

A tree, rarely more than 15°—20° high, with a well-developed trunk 5′—6′ in diameter, stout spreading branches forming a broad low flat-topped head, and stout branchlets orange-brown in their first season, becoming dark gray-brown the following year, and armed with thick straight or slightly curved bright chestnut-brown shining spines 1′—2′ in length.

Distribution. Low limestone ridges opposite Lachine near the south bank of the St. Lawrence River, and on the Island of Montreal, Province of Quebec; near Cornwall, Ontario.

107. Cratægus hudsonica Sarg.

Leaves ovate or slightly obovate, acute, gradually and abruptly narrowed and mostly concave-cuneate at the entire base, sharply and often doubly serrate above with straight or incurved glandular teeth, and frequently slightly divided above the middle into short acute lobes, nearly fully grown when the flowers open at the end of May, and then thin, light yellow-green, smooth and glabrous above with the exception of a few short white scattered hairs on the midrib, and pale and glabrous below, and at maturity thin and firm in texture, glabrous, 2′—2½′ long, and 1½′—1¾′ wide, with a slender yellow midrib, and 5 or 6 pairs of thin primary veins extending obliquely to the point of the lobes; petioles slender, wing-margined above, glandular, at first slightly hairy, becoming glabrous and rose color toward the base, ¾′—1′ in length; leaves at the end of vigorous shoots broad-ovate to suborbicular, full and rounded or broad-cuneate at the wide base, deeply divided into broad lateral lobes, and 2′—3′ long and wide. Flowers about ¾′ in diameter, on long slender pedicels, in broad usually 10—12-flowered glabrous corymbs; calyx-tube narrowly obconic, glabrous, the lobes gradually narrowed from a broad base, acuminate, glandular-serrate often only below the middle, glabrous on the outer surface, slightly hairy on the inner surface; stamens 20; anthers rose color; styles 3—5. Fruit ripening early in September, in few-fruited drooping clusters, subglobose, crimson, pruinose, marked by numerous pale dots, about ⅝′ in diameter; calyx enlarged, with a deep broad cavity, and closely appressed serrate lobes villose on the upper side; flesh thick, yellow, dry and mealy; nutlets 3—5, rounded at base and narrowed and rounded at apex, rounded and sometimes ridged on the back with a high rounded ridge, about /16′ long.

A tree, sometimes 20° high, with a tall trunk 8′—10′ in diameter, covered with pale scaly bark, heavy ascending and spreading branches forming a broad open head, and stout ascending glabrous branchlets dark orange color when they first appear and light orange-brown and lustrous during their first winter, and armed with numerous slender straight or slightly curved bright red-brown shining spines 1½′—2′ long; sometimes a broad bush, with numerous stout spreading stems.

Distribution. Rolling hills in the valley of the Hudson River, near Albany, Albany County, New York.

108. Cratægus coccinioides Ashe.

Leaves broad-ovate, acute, full and rounded or truncate at base, sharply and often doubly serrate with straight glandular teeth, and divided above the middle into short acute lobes, as they unfold conspicuously plicate, very lustrous, yellow-green, and villose on the lower side of the midrib with a few short pale hairs usually persistent during the season, about half grown when the flowers open early in May, and at maturity thin and firm in texture, rather rigid, dull dark green and smooth on the upper surface, pale on the lower surface, 2½′—3′ long, and 2′—2½′ wide, with a thin pale yellow midrib deeply impressed above and often bright red toward the base after midsummer, and slender primary veins arching to the point of the lobes; turning late in October gradually bright orange and scarlet; petioles glandular on the upper side with minute-stalked dark red glands, at first villose, soon glabrous, often bright red or pink toward the base, ¾′—1′ in length; leaves at the end of vigorous shoots more or less cordate at base and usually 3½′—4′ long and wide. Flowers ¾′ in diameter, in very compact 5—7-flowered glabrous or slightly villose corymbs, with coarsely serrate oblong-obovate acute bracts and bractlets, conspicuously glandular with large bright red glands; calyx-tube broadly obconic, glabrous, the lobes gradually narrowed from a broad base, acute and coarsely glandular-serrate; stamens 20; anthers large, deep rose color; styles 5, surrounded at base by a ring of pale tomentum. Fruit ripening early in October and falling gradually during a month or six weeks, on stout pedicels, in few-fruited compact erect clusters, subglobose, much flattened at the ends, often obscurely angled, dark crimson, very lustrous, marked by numerous large pale dots, ¾′ long, and ⅞′ wide; calyx much enlarged and conspicuous, with spreading or erect lobes bright red on the upper side near the base; flesh thick, firm, subacid, more or less deeply tinged with red; nutlets 5, comparatively small, light-colored, narrow at the ends, acute at apex, rounded at base, rounded and slightly ridged on the back, about ⅓′ long.

A tree, sometimes 20° high, with a stem 8′—10′ in diameter, covered with dark brown scaly bark, stout spreading light gray branches forming a broad handsome head, and stout nearly straight glabrous bright chestnut-brown very lustrous branchlets armed with thick dark reddish purple shining spines 1½′—2′ long.

Distribution. Dry woods in the neighborhood of St. Louis, Missouri; in eastern Kansas.

XI. Rotundifoliæ.

Coccineæ Sarg.

Conspectus of the Arborescent Species.

Stamens 10; leaves coriaceous.

Leaves elliptic or obovate; fruit subglobose, dark crimson; anthers pale yellow.

109. C. rotundifolia (A).

Leaves elliptic or ovate; fruit short-oblong to oblong-obovoid, bright carmine-red; anthers rose color.

110. C. Jonesæ (A).

Stamens 20; leaves subcoriaceous, rhombic to oblong-obovate; fruit short-oblong to subglobose, dark dull red or rusty orange-red; anthers pale yellow.

111. C. Margaretta (A, C).

109. Cratægus rotundifolia Moen.

Cratægus coccinea var. rotundifolia Sarg.

Leaves elliptic or obovate, acute or acuminate, gradually narrowed from above the middle to the cuneate entire base, finely and often doubly serrate above with incurved or straight teeth tipped with minute dark glands, and divided above the middle into several short acute lateral lobes, about half grown when the flowers open at the end of May, and then thin, light yellow-green and glabrous, and at maturity coriaceous, dark green, smooth and lustrous on the upper surface, paler on the lower surface, 1½′—2′ long, and 1′—1½′ wide, with a thin midrib, and 4 or 5 pairs of primary veins extending to the point of the lobes; petioles slender, glandular, slightly winged at apex, glabrous, often dark red toward the base, ¾′—1′ in length; leaves at the end of vigorous shoots oblong-ovate, oval or often nearly orbicular, more deeply lobed, and frequently 2½′—3′ long and wide. Flowers ½′—¾′ in diameter, on slender pedicels, in broad loose many-flowered glabrous corymbs; calyx-tube broadly obconic, glabrous, the lobes gradually narrowed from a broad base, acute, coarsely glandular-serrate, glabrous, often bright red toward the apex; stamens 10; anthers small, pale yellow; styles 3 or 4. Fruit ripening and falling late in October, on short stout pedicels, in drooping many-fruited glabrous clusters, subglobose but occasionally rather longer than broad, dark crimson, marked by scattered dark dots, about ½′ in diameter; calyx enlarged, conspicuous, the lobes bright red on the upper side toward the base, wide-spreading or erect; flesh thin, yellow, dry and sweet; nutlets 3 or 4, rounded at the ends, about ¼′ long.

A bushy tree, occasionally 20° high, with a short trunk 8′—10′ in diameter, covered with dark red-brown scaly bark, stout ascending branches forming a broad round-topped symmetrical head, and slender glabrous branchlets light green when they first appear, bright red-brown and lustrous during their first year, and ultimately ashy gray, and armed with many stout straight or slightly curved chestnut-brown shining spines 1′—1½′ long.

Distribution. Nova Scotia, southern Quebec and Ontario to Manitoba and Saskatchewan (Saskatoon), and southward through New England, eastern and northern New York, the southern peninsula of Michigan and northern Indiana; in Pennsylvania (Lackawanna, Bucks, Northampton and Blair Counties); common in the New England coast region; a form (var. pubera Sarg.) with young leaves covered above with soft pale hairs and pubescent on the under side of the midrib and veins and villose petioles, flowers with a pubescent calyx-tube, in villose corymbs, becoming pilose when the fruit ripens, and young branchlets covered with long matted pale hairs, ranges from Newfoundland to the shores of Lake St. John, Province of Quebec, northern Ontario, Winnipeg and Manitoba, and southward through the maritime provinces of Canada, New England to southern Connecticut, northern and western New York (near Buffalo, Essex County), the northern peninsula of Michigan, northeastern Wisconsin; in central Minnesota (St. Cloud, Stearns County); common northward.

110. Cratægus Jonesæ Sarg.

Leaves elliptic to ovate, acute, gradually narrowed or broad-cuneate at the entire base, coarsely doubly serrate above with spreading or incurved teeth tipped with deciduous dark red glands, and usually divided above the middle into 2 or 3 pairs of short acute or acuminate lobes, more than half grown when the flowers open during the first week of June, and then membranaceous and coated with soft pale hairs most abundant on the under side of the midrib and principal veins, and at maturity thick and coriaceous, dark green and lustrous on the upper surface, pale and puberulous on the lower surface, 3′—4′ long and 2′—3′ wide, with a stout midrib, 4—6 pairs of primary veins and conspicuous secondary veinlets; petioles stout, more or less winged toward the apex, villose, ultimately glabrous, tinged with red below the middle, 1½′—2′ in length, after midsummer often twisted at base, bringing the lower surface of the leaf to the light; leaves at the end of vigorous shoots usually more coarsely serrate and much more deeply lobed, with broadly winged petioles, and falcate coarsely glandular-serrate stipules sometimes 1′ in length. Flowers 1′ in diameter, on long slender pedicels, in broad loose lax many-flowered tomentose corymbs; calyx-tube narrowly obconic, tomentose, the lobes abruptly narrowed from a broad base, long, acute, entire, villose; stamens 10; anthers large, rose color; styles 2, or generally 3, surrounded at base by a narrow ring of pale tomentum. Fruit ripening usually early in October, on slender elongated pedicels, in broad many-fruited drooping glabrous or puberulous clusters, short-oblong to oblong-obovoid, rounded at the ends, bright carmine-red, marked by occasional large dots, ¾′—1′ long, and ¾′ in diameter; calyx conspicuous, with enlarged and elongated closely appressed lobes; flesh thick, yellow, sweet and mealy; nutlets 3 or rarely 2, thick, narrowed and acute at base, full and broad at apex, rounded and ridged on the back with a high broad ridge, about /16′ long.

A tree, sometimes 20° high, with a tall trunk often a foot in diameter, covered with dark brown scaly bark, ascending or spreading branches forming a broad open irregular head, and stout branchlets tomentose early in the season, becoming orange-brown, glabrous and very lustrous during their first summer, and light gray the following year, and armed with stout straight or curved chestnut-brown shining spines 2′—3′ long and usually pointed toward the base of the branch.

Distribution. Rocky shores of sounds and bays; coast of Maine, Islesboro and Belfast Bay to the island of Mount Desert (Waldo and Hancock Counties); in hedges, near Fredericton, York County, New Brunswick; Rivière du Loup, Kamouraska County, Province of Quebec (Brother Victorin).

111. Cratægus Margaretta Ashe.

Leaves broad-rhombic, oblong-obovate or rarely ovate, acute or rounded at apex, gradually narrowed and usually entire below, coarsely often doubly crenately-serrate above with usually glandless teeth, and divided above the middle or frequently only at apex into short broad rounded or acute lobes when the flowers open in May, thin and roughened above by short pale hairs and glabrous below, and at maturity firm and rather leathery in texture, or subcoriaceous, glabrous, smooth, dark green and somewhat lustrous on the upper surface, pale on the lower surface, 1′—1¼′ long, and 1′ wide, with a yellow midrib, and 3—5 pairs of primary veins extending obliquely to the point of the lobes; petioles slender, often slightly winged toward the apex, glandular at first with minute dark red caducous glands, ½′—1′ in length; leaves at the end of vigorous shoots broad-ovate or semiorbicular, usually more deeply and more generally lobed, often 3′ long and 2′—3′ wide. Flowers about ¾′ in diameter, on long slender pedicels, in 3—12-flowered thin-branched slightly villose corymbs; calyx-tube narrowly obconic, slightly villose toward the base, or glabrous, the lobes gradually narrowed from below, acuminate or short-pointed at apex, finely and irregularly glandular-serrate, glabrous or villose on the inner surface; stamens usually 20; anthers small, light yellow; styles 2 or 3, surrounded at the base by a narrow ring of pale tomentum, and villose below the middle with occasional long spreading hairs. Fruit ripening and falling at the end of September, in few-fruited drooping clusters, short-oblong, rounded at the ends, or subglobose and flattened at the ends, dull dark red or rusty orange-red or rarely yellow, marked by occasional dark dots, and about ½′ long; calyx only slightly enlarged, the lobes spreading or erect and frequently deciduous before the fruit ripens; flesh thin, yellow, dry and mealy; nutlets 2 or 3, broad and rounded at base, acute at apex, conspicuously grooved and ridged on the back with a broad rounded ridge, about ¼′ long.

A tree, occasionally 25° high, with a straight trunk 4′—6′ in diameter, covered with thin dark gray-brown bark, small rather erect branches forming a narrow open head, and slender branchlets, orange-green, glabrous or sometimes pubescent when they first appear, becoming bright chestnut-brown and lustrous, and ashy gray or gray tinged with red during their second year, and armed with thin straight or slightly curved bright chestnut-brown spines ¾′—1½′ long.

Distribution. Central Iowa (Steamboat Rock, Harden County, Cedar Rapids, Linn County), southward to Missouri (Hannibal, Marion County, Webster, St. Louis County to the neighborhood of Springfield, Greene County), and eastward to northeastern Illinois (Downers Grove, Dupage County); through north central Indiana to southern Michigan (Kalamazoo and Ingham Counties); through central and southern Ohio to the southeastern part of the state (Washington County); southeastern Ontario (London and Oakwood); in central Tennessee (West Nashville, Davidson County).

XII. Intricatæ.

Conspectus of the Arborescent Species.

Stamens 10; leaves broad-ovate to oval.

Fruit depressed-globose, yellow-green flushed with russet-red; anthers pale yellow; calyx-lobes eglandular.

112. C. Boyntonii (A, C).

Fruit subglobose, red or russet-red; anthers pale rose color; calyx-lobes glandular with stalked glands.

113. C. Buckleyi (A).

Stamens 20.

Leaves oval to ovate or oblong-obovate; fruit short-oblong, dull red, often with a bright russet face; stamens usually 5—15; anthers small, pale yellow.

114. C. venusta (C).

Leaves oblong-ovate to elliptic or ovate; fruit subglobose to short-oblong, yellow or orange-yellow, more or less flushed with red; anthers large, purple.

115. C. Sargentii (C).

112. Cratægus Boyntonii Beadl.

Leaves broad-ovate to oval, acute, rounded or cuneate at the entire glandular base, sharply and often doubly serrate above with glandular teeth, and frequently divided into 2 or 3 pairs of short broad acute lateral lobes, when they unfold deep bronze-red, slightly glandular and viscid, nearly fully grown when the flowers open early in May, and then membranaceous and glabrous or occasionally slightly pilose, and at maturity subcoriaceous, glabrous, yellow-green on the upper surface, pale on the lower surface, 1′—2½′ long, and 1′—2′ wide, with a thin pale yellow midrib and 4—7 pairs of slender veins; petioles stout, glandular often to the base with bright red glands, slightly winged above, usually about ½′ in length; leaves at the end of vigorous shoots often as broad as long, truncate or cordate at base, and more coarsely dentate and more deeply lobed. Flowers about ¾′ in diameter, on short slender pedicels, in compact 4—10-flowered compound corymbs; calyx-tube broadly obconic, the lobes abruptly narrowed from a broad base, acute or rounded at apex, entire or obscurely and irregularly glandular-serrate above the middle; stamens 10; anthers large, pale yellow; styles 3—5, surrounded at base by a broad thick ring of hoary tomentum. Fruit ripening and falling early in October, on short stout pedicels, in few-fruited erect clusters, depressed-globose, more or less angled, yellow-green flushed with russet-red, marked with small dark dots, usually about ½′ in diameter; calyx prominent, the large spreading lobes often deciduous before the fruit ripens; nutlets 3—5, acute or acuminate at apex, rounded at the narrow base, about ¼′ long.

A tree, occasionally 20° high, with a tall straight trunk 6′—8′ in diameter, sometimes armed with long gray compound spines, stout ascending branches forming a narrow open irregular or occasionally a round-topped head, and glabrous branchlets furnished with many thin nearly straight light chestnut-brown spines 1½′—2′ long; or more often a shrub, with numerous stems.

Distribution. Banks of streams, the borders of fields and upland woods in the southern Appalachian foothill region from southern Virginia to northern Georgia; in northern Alabama, southeastern Kentucky, and eastern Tennessee; sometimes ascending to altitudes of 3000° above the sea.

113. Cratægus Buckleyi Beadl.

Leaves broad-ovate or oval, acute, rounded or subcordate or narrowed and concave-cuneate at the entire base, coarsely often doubly serrate above with straight glandular teeth, and more or less incisely lobed with acuminate lateral lobes, more than half grown when the flowers open about the middle of May and then pale green and glabrous with the exception of a few caducous hairs on the upper side of the base of the midrib, and at maturity dark green above, paler below, 1½′—2′ long, and 1½′—2′ wide; petioles stout, conspicuously glandular above the base, wing-margined at the apex, glabrous, ½′—¾′ in length. Flowers about ¾′ in diameter, on slender glabrous pedicels, in compact 3—7-flowered simple corymbs, with conspicuously glandular bracts and bractlets; calyx-tube broadly obconic, glabrous, the lobes broad, acuminate, laciniately cut toward the apex, and glandular with stipitate glands; stamens 10; anthers pale rose color; styles 3—5, surrounded at base by tufts of pale hairs. Fruit ripening late in September or in October, subglobose, usually angled, red or russet-red, about ½′ in diameter; calyx little enlarged, with spreading or reflexed lobes; flesh thin, dry and mealy; nutlets 3—5, broad and rounded at base, rounded at the slightly narrowed apex, prominently ridged on the back, with a broad grooved ridge, about /16′ long.

A tree, often 25° high, with a trunk 4′—7′ in diameter and sometimes 10°—12° long, covered with gray or often dark brown scaly bark, stout spreading or ascending branches, and thick glabrous red-brown branchlets armed with thin straight shining spines ½′ long, becoming much longer and branched on the trunk and large branches.

Distribution. Southwestern Virginia, through western North Carolina to eastern Tennessee; usually at altitudes between 2000° and 3000°; common on wooded slopes with Oaks, Hickories, and Pines.

114. Cratægus venusta Beadl.

Leaves oval to ovate or occasionally to oblong-ovate, acute, gradually or abruptly narrowed and cuneate or rounded at the entire base, finely serrate above with usually incurved glandular teeth, and frequently slightly and irregularly divided above the middle into 1—3 pairs of short broad acute lobes, when they unfold dark bronze color, with a few scattered pale caducous hairs on the upper surface, about half grown when the flowers open from the 20th to the end of April, and then yellow-green, smooth and glabrous, and at maturity dark dull green above, pale below, 2½′ long, and 1½′ wide, with a stout midrib and 4—7 pairs of thin primary veins; late in the autumn turning, especially those on leading shoots deep orange or scarlet; petioles stout, glandular, more or less winged above, ½′—¾′ in length; leaves at the end of vigorous shoots generally broad-ovate, rounded at base, deeply lobed with broad lobes, and often 3½′ long and 3′ wide. Flowers 1′ in diameter, on short pedicels, in 4—9-flowered compact corymbs, their bracts and bractlets like the inner bud-scales coarsely glandular-serrate and bright red before falling; calyx-tube broadly obconic, the lobes gradually narrowed from a broad base, acute, coarsely glandular-serrate often only below the middle; stamens 15—20, usually 15—17; anthers small, pale yellow; styles 3—5, surrounded at the base by a ring of pale hairs. Fruit ripening and falling from the 1st to the middle of October, on stout pedicels often 1′ long, in few-fruited clusters, short-oblong, rounded at the ends, dull red, often with a bright russet face, and marked by occasional large dark dots; calyx prominent, with a long tube, and spreading lobes often deciduous before the fruit ripens; flesh thick, yellow, dry and mealy; nutlets 3—5, narrow and acute at base, broad, about ¼′ long.

A bushy tree, often 25° high, with a short trunk a foot in diameter, furnished like the large branches with innumerable stout much-branched spines frequently 6′ long, and slender nearly straight glabrous dark chestnut-brown branchlets, armed with many stout straight or slightly curved dark chestnut-brown shining spines frequently pointing toward the base of the branch, and 1½′—2½′ long.

Distribution. Open Oak and Hickory-woods on the dry slopes of Red Mountain in the southern part of the city of Birmingham, Jefferson County, Alabama.

115. Cratægus Sargentii Beadl.

Leaves oblong-ovate to elliptic or rarely to ovate, acute or acuminate at apex, gradually or abruptly cuneate or rounded at the nearly entire base, irregularly doubly serrate above with straight or incurved glandular teeth, and usually irregularly divided into 3 or 4 pairs of short broad acute or acuminate lobes, nearly fully grown when the flowers open late in April, and then subcoriaceous, pale yellow-green, and villose on the midrib with scattered pale caducous hairs, and at maturity lustrous, dark yellow-green above, pale below, 2′—3′ long, and 1½′—2′ wide, with a thin midrib, 5—7 pairs of thin light yellow veins and conspicuous reticulate veinlets; turning in the autumn bright yellow and red; petioles slender, glandular, more or less broadly winged toward the apex, ½′—¾′ in length; leaves at the end of vigorous shoots oblong-ovate, concave-cuneate at base, often 3′ long and 2′ wide, their petioles broadly wing-margined to below the middle. Flowers nearly 1′ in diameter, on long thin slightly villose pedicels, in 2—5 usually 3-flowered simple corymbs, with coarsely glandular-serrate bracts and bractlets; calyx-tube narrowly obconic, glabrous or slightly villose, the lobes foliaceous, acute, coarsely glandular-serrate above the middle; stamens 20; anthers large, dark rose color; styles 3—5, usually 4, surrounded at base by a narrow ring of pale hairs. Fruit ripening and falling about the middle of September, often only a single fruit maturing from a flower-cluster, subglobose to short-oblong, rounded at the ends, yellow or orange-yellow, generally more or less flushed with red, marked by occasional large dark dots, ⅓′—½′ long; calyx prominent, with an elongated tube and closely appressed lobes; flesh yellow, thin and hard; nutlets 3—5, usually 4, about ¼′ long.

An intricately branched tree, rarely more than 20° high, with a tall trunk 6′—7′ in diameter, stout ascending branches forming a narrow or sometimes a round flat-topped head, and glabrous branchlets armed with thin straight or slightly curved dark chestnut-brown shining spines, ¾′—1½′ long; often a large shrub, with few or many stems.

Distribution. Rocky woods and bluffs in the foothill region of northwestern Georgia (cliffs of the Coosa River near Rome, Floyd County), southeastern Tennessee (near Chattanooga, Hamilton County, and Tracy City, Grundy County), and northeastern Alabama; very abundant in Alabama at Valley Head, De Kalb County, and on the low ridges extending southward to the neighborhood of Birmingham, Jefferson County.

XIII. Pulcherrimæ.

Conspectus of the Arborescent Species.

Leaves oval to ovate or nearly orbicular, their lobes acute or rounded; fruit bright red.

116. C. opima (C).

Leaves ovate to oval or obovate, their lobes acute; fruit orange-red.

117. C. robur (C).

116. Cratægus opima Beadl.

Leaves oval to ovate or nearly orbicular, acute, gradually or abruptly narrowed and cuneate at the entire base, finely serrate above with incurved teeth, and usually divided above the middle into short acute, acuminate or rounded lobes, half grown when the flowers open the middle of April, and then glabrous with the exception of a few short caducous hairs on the midrib and veins, and at maturity light green on the upper surface, pale on the lower surface, 1½′ long, and 1¼′ wide, with a slender midrib, and 5 or 6 pairs of arcuate primary veins spreading to the point of the lobes; petioles narrowly winged at the apex, usually about ¾′ in length; leaves at the end of vigorous shoots sometimes rounded or nearly truncate at base and 1½′—2½′ long and broad. Flowers about ⅔′ in diameter, on short slender pedicels, in compact few-flowered glabrous corymbs; calyx-tube broadly obconic, glabrous, the lobes gradually narrowed from a broad base, acute, entire or sparingly glandular-serrate, tipped with dark red glands, glabrous on the outer surface, puberulous on the inner surface; stamens 20; anthers dark rose color; styles 3—5, surrounded at base by a narrow ring of snowy white tomentum. Fruit ripening about the 1st of October and then remaining on the branches for several weeks, on short stout pedicels, in compact few-fruited erect or drooping clusters, subglobose, often rather longer than broad, bright red, about ¼′ in diameter; calyx prominent, with a well-developed tube, and much enlarged closely appressed lobes often deciduous with the tube before the fruit becomes entirely ripe; flesh thin, yellow, dry and mealy; nutlets 3—5, thin, ⅛′ long.

A tree, 20°—25° high, with a tall, slender often spiny trunk covered with ashy gray bark nearly black at the base of old trees, spreading and ascending branches forming a rounded or oval usually open head, and thin nearly straight bright red-brown glabrous branchlets becoming gray tinged with red or brown in their second season, and armed with thin nearly straight bright chestnut-brown lustrous spines, 1′—1½′ long.

Distribution. Open woods in clay soil in the neighborhood of Greenville, Butler County, Alabama; common near Tallahassee, Leon County, Florida.

117. Cratægus robur Beadl.

Leaves ovate, oval or obovate, acute or acuminate, entire or sparingly glandular below, finely serrate above with incurved glandular teeth, and incisely lobed above the middle with numerous short acute lobes, nearly fully grown when the flowers open at the end of March, and then membranaceous and dark yellow-green and lustrous, and at maturity yellow-green, 1½′—2′ long, and 1′—1½′ wide, with a slender yellow midrib, and thin primary veins extending very obliquely to the point of the lobes; turning in the autumn orange, yellow, or brown; petioles slender, slightly wing-margined toward the apex, sparingly glandular, ½′—1′ in length; leaves at the end of vigorous shoots broadly ovate, cuneate or nearly truncate at the wide base, deeply divided into broad lateral lobes, often 2′—3′ long and broad, with a stout broadly winged petiole frequently 1′ long. Flowers 1⅛′—1¼′ in diameter, on long slender pedicels, in 5—10-flowered glabrous corymbs, with large conspicuously glandular bracts and bractlets; calyx-tube narrowly obconic, glabrous, the lobes gradually narrowed from a broad base, glabrous, entire or sparingly serrate; stamens 20; anthers pale rose color; styles 3—5, surrounded at base by a narrow ring of pale hairs. Fruit ripening in September and October, on elongated, slender pedicels, in few-fruited drooping clusters, subglobose, orange-red, about ½′ in diameter; calyx-lobes deciduous before the maturity of the fruit leaving a narrow ring round the shallow cavity; flesh thin and firm; nutlets 3—5, broad, rounded at the ends, barely grooved on the rounded back, /16′ long and nearly as broad.

A tree, 20°—25° high, with a trunk 4′—6′ in diameter, covered with gray or brown scaly bark, spreading or ascending branches, and slender red-brown branchlets unarmed or armed with stout spines ¾′—1′ long; more often a large much-branched shrub, with one or more stems.

Distribution. Woods and borders of fields, northwestern Florida; common in the neighborhood of Tallahassee, Leon County.

XIV. Bracteatæ.

Conspectus of the Arborescent Species.

Leaves oval to broad-obovate, subcoriaceous; corymbs many-flowered; stamens 10—20, usually 20; fruit bright red or orange-red.

118. C. Harbisonii (C).

Leaves broad-ovate or rarely obovate, thin; corymbs 3—10-flowered; stamens 20; fruit bright red.

119. C. Ashei (C).

118. Cratægus Harbisonii Beadl.

Leaves oval to broad-obovate, acute at apex, cuneate or rounded at the entire base, and coarsely serrate above with straight glandular teeth, when they unfold roughened above by stout, rigid pale hairs, and soft and pubescent below, nearly fully grown early in May when the flowers open, and then thin, dark yellow-green above and pale below, and at maturity subcoriaceous; pale on the lower surface, 2′—2½′ long, and 1′—1½′ wide, with a stout midrib and primary veins deeply impressed on the upper side of the leaf, and conspicuous reticulate veinlets; petioles stout, villose, more or less winged above, ¼′—½′ in length; leaves at the end of vigorous shoots broad-ovate, cuneate and decurrent on their stouter petiole, 3′—4′ long, and 2½′—3′ wide, with lunate coarsely glandular-dentate stipules frequently ½′ long. Flowers ¾′ in diameter, in broad loose usually 10—12-flowered corymbs, with broad acute conspicuous glandular-serrate bracts and bractlets; calyx-tube broadly obconic, densely villose at the base and glabrous or pubescent above, the lobes elongated, gradually narrowed from a broad base, acute, bright green, more or less hairy, coarsely glandular-serrate, with large stipitate dark red glands; stamens 10—20, usually 20; anthers large, light yellow; styles 3—5. Fruit ripening and falling early in October, subglobose, often rather longer than broad, bright red or orange-red, marked by numerous large dark dots; calyx enlarged, with spreading glandular lobes often deciduous before the fruit ripens; flesh yellow, thick, dry and mealy; nutlets 3—5, narrowed at the ends, ¼′ long.

A tree, sometimes 25° high, with a trunk 10′—12′ in diameter, covered with light gray or gray-brown bark, and often armed with straight or much-branched spines, wide-spreading light gray or reddish branches forming a rather open symmetrical head, and slender branchlets coated when they first appear with long spreading white hairs, pubescent or glabrous and light red-brown or orange-brown during their first season, becoming dark or light gray the following year, and furnished with numerous usually stout straight dark reddish brown shining spines 1½′—2′ long.

Distribution. Dry limestone hills and ridges; West Nashville, Davidson County, Tennessee; common.

119. Cratægus Ashei Beadl.

Leaves broad-ovate or occasionally obovate, acute and generally short-pointed at apex, gradually or abruptly narrowed and cuneate and usually entire at base, coarsely and occasionally doubly serrate above with straight or incurved teeth tipped with small dark glands, when they unfold roughened on the upper surface with short pale hairs and pubescent below, nearly fully grown and membranaceous when the flowers open early in May, and at maturity thin but firm in texture, pale and puberulous on the lower surface on the slender midrib and primary veins, about 2′ long and 1½′ wide; petioles stout, broadly winged above, glandular, pubescent early in the season but ultimately nearly glabrous, about ½′ in length; leaves at the end of vigorous shoots usually broadly oval or nearly orbicular, rounded or short-pointed at apex, 2½′—3′ long, and 2′—2½′ wide. Flowers ¾′ in diameter, on slender hairy pedicels, in 3—10-flowered simple or compound corymbs, with broad conspicuous glandular bracts and bractlets; calyx-tube broadly obconic, thickly coated with long matted reflexed white hairs, the lobes broad, acute, nearly glabrous on the outer surface, villose on the inner surface, glandular with small stout stipitate glands; stamens 20; anthers small, yellow; styles 3—5, surrounded at base by a narrow ring of pale hairs. Fruit ripening and falling late in September or early in October, on stout villose or glabrous pedicels, in few-fruited clusters, subglobose or rather longer than broad, bright red, marked by large scattered dots, more or less villose toward the ends, about 1′ in diameter; calyx conspicuous, with elongated coarsely glandular-serrate, erect incurved or reflexed lobes; flesh thick and yellow; nutlets 3—5, thin, acute at the ends, ⅓′ long.

A tree, rarely more than 20° high, with a slender trunk covered with smooth light gray or red-brown bark becoming fissured and scaly on old individuals, stout ascending branches forming a pyramidal or oval head, and slender branchlets coated when they first appear with long pale matted reflexed hairs, soon becoming nearly glabrous, lustrous, orange-brown or reddish brown, and light gray or gray tinged with red during their second season, and armed with straight or slightly curved thin dark red-brown shining spines 1′—1½′ long.

Distribution. Abandoned fields, and woods, growing usually on clay soils; near Montgomery, Montgomery County, and Gallion, Hale County, Alabama.

XV. Flavæ.

Conspectus of the Arborescent Species.

Stamens 20.

Anthers pink or purple.

Corymbs usually 3—6-flowered.

Leaves elliptic to broad-ovate, yellow-green; fruit dark orange-brown.

120. C. flava (C).

Leaves ovate to obovate or orbicular, bright yellow-green; fruit obovoid, dark orange color with a red cheek.

121. C. visenda (C).

Leaves obovate or ovate, dark green; fruit subglobose to short-oblong, red or orange-red.

122. C. ignava (C).

Corymbs 1—5-flowered.

Leaves broad-obovate to nearly orbicular, bright green; fruit globose or depressed-globose, bright red.

123. C. consanguinea.

Leaves obovate, bright green; fruit ellipsoidal to short-oblong, orange-red; anthers pink.

124. C. tristis.

Anthers yellow (doubtful in 128, 133).

Leaves yellow-green.

Leaves 3-nerved.

Leaves obovate-cuneate, often 3-lobed at apex; fruit obovoid to subglobose, bright orange-red and lustrous; corymbs tomentose.

125. C. floridana.

Leaves obovate; fruit subglobose to short-oblong, dull brownish yellow; corymbs glabrous.

126. C. lacrimata.

Leaves with numerous primary veins.

Leaves thin.

Leaves scabrate above at maturity, obovate, rounded or abruptly short-pointed at apex; fruit subglobose to short-oblong, bright orange-red.

127. C. Ravenelii (C).

Leaves smooth above at maturity.

Leaves obovate to obovate-cuneiform; fruit subglobose, bright red.

128. C. senta (A).

Leaves obovate to oval or orbicular; fruit subglobose to ellipsoidal, orange-red or red and orange.

129. C. annosa (C).

Leaves subcoriaceous.

Flowers in 3—5-flowered corymbs.

Leaves obovate; fruit globose or depressed-globose, orange-yellow with a red cheek.

130. C. panda (C).

Leaves obovate to oblong-ovate, minutely serrate; fruit globose, red or yellow.

131. C. integra (C).

Flowers in 1 or 2-flowered corymbs; leaves spathulate; fruit obovoid, red.

132. C. recurva (C).

Leaves conspicuously blue-green, broad-ovate to orbicular; fruit subglobose to short-oblong, light red, puberulous at the ends.

133. C. dispar (C).

Stamens 10; anthers yellow; leaves broad-obovate to oval or rhombic, dark yellow-green; fruit subglobose, dull orange-red, often slightly villose at the ends.

134. C. aprica (C).

120. Cratægus flava Ait.

Leaves elliptic to broad-obovate, acute or rarely rounded at apex, gradually narrowed and cuneate at the glandular base, and coarsely doubly serrate above with broad straight or incurved teeth tipped with large dark red stipitate glands, when they unfold bronze color, villose above with short pale caducous hairs most abundant near the base of the midrib and pubescent below on the midrib and veins, about half grown when the flowers open from the 10th to the 20th of April, and at maturity membranaceous, yellow-green, usually about 2′ long and 1½′ wide, with a slender yellow midrib and 3 or 4 pairs of primary veins usually puberulous on the under side and only slightly impressed above; petioles slender, glandular, winged nearly to the base, generally more or less villose, after midsummer often light red on the lower side, and about ½′ in length; leaves at the end of vigorous shoots frequently 3′ long and 2′ wide, and sometimes broad-ovate, 3-lobed or divided into 2 or 3 pairs of lateral lobes, their petioles 1′—1½′ long, broadly winged and conspicuously glandular, and foliaceous lunate or elliptic coarsely glandular-serrate stipules. Flowers about ¾′ in diameter, on short slender pedicels, in few-flowered simple or compound slightly villose compact corymbs, with lanceolate acute coarsely glandular-serrate bracts and bractlets; calyx-tube broadly obconic, glabrous, the lobes wide, acute, usually laciniately divided, very glandular; stamens 20; anthers large, dark rose color. Fruit ripening early in October and soon falling, in few-fruited drooping clusters, short-oblong, full and rounded at the ends, dark orange-brown, ½′—⅝′ long, and ⅓′—½′ in diameter; calyx prominent, with a long narrow tube, and enlarged closely appressed lobes often deciduous before the fruit ripens; flesh thick, orange color, dry and mealy; nutlets 5, gradually narrowed and rounded at the ends, ridged and deeply grooved on the back with a high narrow ridge, about ½′ long.

A tree, 15°—20° high, with a tall trunk 8′—10′ in diameter, covered with thin dark brown bark tinged with red and divided into narrow rounded ridges, stout ascending branches forming an open and somewhat irregular head sometimes 20° across, and slender slightly zigzag glabrous branchlets dark green deeply tinged with red when they first appear, becoming dull red-brown or orange-brown during their first season, darker the following year, and ultimately dark gray-brown, and armed with thin nearly straight bright chestnut-brown spines ¾′—1¼′ long.

Distribution. Dry sandy soil on the sand hills of Summerville, near Augusta, Richmond County, Georgia, and at River Junction, Gadsden County, Florida.

121. Cratægus visenda Beadl.

Leaves ovate, obovate, or orbicular, short-pointed and acute or occasionally broad and rounded at apex, concave-cuneate and gradually narrowed at the mostly entire base, finely serrate above with rounded teeth, glandular with bright red glands, and divided above the middle into short acute lobes, nearly fully grown when the flowers open at the end of March, and then glabrous with the exception of a few short pale hairs on the two surfaces near the base of the midrib, and at maturity thin and firm in texture, bright yellow-green and lustrous above, pale below, glabrous, 1′—1½′ long, and ¾′—1′ wide, with a slender midrib, and thin primary veins extending very obliquely to the point of the lobes; turning yellow, orange, or brown in the autumn; petioles slender, broadly wing-margined above the middle, conspicuously glandular, sparingly villose early in the season, becoming nearly glabrous, ½′—¾′ in length. Flowers about ¾′ in diameter, on short villose pedicels, in simple 3—6-flowered corymbs; calyx-tube broadly obconic, hairy near the base with scattered pale hairs, glabrous above, the lobes broad, acuminate, glandular-serrate, glabrous on the outer, pilose on the inner surface; stamens 20; anthers pale purple; styles 3—5, surrounded at base by small tufts of white hairs. Fruit ripening and falling late in August and early in September, on stout pedicels, usually in 1 or 2-fruited clusters, obovoid, dark orange-colored, with a red cheek, ½′—⅝′ long, nearly ½′ in diameter; calyx enlarged, the lobes coarsely glandular-serrate, puberulous on the upper surface, closely appressed; flesh soft and yellow; nutlets 3—5, obtuse and rounded at the ends, rounded and slightly ridged on the back, about ⅜′ long.

A tree, sometimes 30° high, with a trunk 10′—12′ in diameter, covered with dark gray or brownish bark, crooked horizontal or ascending branches forming a broad irregular head, and stout often contorted branchlets villose when they first appear, soon glabrous, dull reddish brown to ashy gray, and armed with slender straight spines ½′—⅝′ long.

Distribution. Sandy soil near Bristol, Liberty County, Florida.

122. Cratægus ignava Beadl.