A text-book of veterinary anatomy Papyras Classics

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Chapter 352

The Eleventh or Spinal Accessory Nerve

The spinal accessory nerve (N. accessorius) is purely motor. It consists of two parts which differ in origin and function.

The medullary part arises from the lateral aspect of the medulla by several rootlets which are behind and in series with those of the vagus (Fig. ). The spinal part arises from the cervical part of the spinal cord by a series of fasciculi which emerge between the dorsal and ventral roots. The bundles unite to form a trunk which is very small at its origin at the fifth segment of the cord, but increases in size when traced toward the brain, since it continually receives accessions of fibers. It passes through the foramen magnum and joins the medullary part. The trunk thus formed sends its medullary fibers to the tenth and ninth nerves and emerges through the foramen lacerum posterius. It then runs backward and downward with the vagus in a fold of the guttural pouch, separates from that nerve, crosses the deep face of the submaxillary gland and the occipital artery, and divides in the recessus atlantis into dorsal and ventral branches.

It is connected by anastomotic branches with the vagus and hypoglossal nerves and the superior cervical ganglion of the sympathetic, and contributes a branch to the pharyngeal plexus.

The dorsal branch (R. dorsalis) (Figs. , , ) receives a twig from the second and third cervical nerves and turns around the atlantal tendon of the splenius under cover of the mastoido-humeralis. It then passes obliquely through the latter muscle and continues backward on the splenius and the cervical part of the serratus, inclines upward across the anterior deep pectoral and the supraspinatus, and enters the deep face of the trapezius, in which it ramifies.

The ventral branch (R. ventralis) (Fig. ) is smaller. It enters the sterno-cephalicus muscle behind the cervical angle of the parotid gland.

The fibers of the spinal part of the accessory arise from the ventro-lateral cells of the ventral gray column of the cord as far back as the fifth cervical segment. The fibers of the medullary part come chiefly from the nucleus ambiguus in common with the motor fibers of the vagus.